Diplodocoidea Marsh, 1884 vide Barrett and
Upchurch, 1994
= "Diplodocoidea" Marsh, 1884 vide Upchurch, 1993
= Atlantosauria Haeckel, 1895
= Diplodocida Haeckel, 1895
Definition- (Diplodocus longus <- Saltasaurus loricatus)
(Wilson, 2005; modified from Wilson and Sereno, 1998)
Comments- Diplodocoidea was first used in Upchurch's (1993) unpublished
thesis, and later used in three papers in the same volume. Upchurch (1994) uses
it in quotes and notes it is only used as a label. Barrett and Upchurch (1994)
state its definition is in Upchurch's thesis, but do not otherwise disclaim
the name. Hunt et al. (1994) use the superfamily later in the volume, but misspell
it Diplodocoidae. Upchurch (1995) is the usual reference given for the name.
References- Marsh, 1884. Principal characters of American Jurassic dinosaurs.
Part VII. Diplodocidae, a new family of the Sauropoda. American Journal of Science.
27, 161-167.
Haeckel, 1895. Systematische Phylogenie der Wirbelthiere: (Vertebrata). 660
pp.
Upchurch, 1993. The anatomy, phylogeny and systematics of sauropod dinosaurs.
PhD thesis. University of Cambridge. 489 pp.
Hunt, Lockley, Lucas and Meyer, 1994. The global sauropod fossil record. Gaia.
10, 261-279.
Barrett and Upchurch, 1994. Feeding mechanisms of Diplodocus. Gaia. 10,
195-203.
Upchurch, 1994. Sauropod phylogeny and palaeoecology. Gaia. 10, 249-260.
Upchurch, 1995. The evolutionary history of sauropod dinosaurs. Philosophical
Transactions of the Royal Society of London, Series B. 349, 365-390.
Wilson and Sereno, 1998. Early evolution and higher-level phylogeny of sauropod
dinosaurs. Society of Vertebrate Paleontology Memoir 5. Journal of Vertebrate
Paleontology. 18(2 suppl), 68 pp.
Wilson, 2005. Overview of sauropod phylogeny and evolution. in Curry Rogers
and Wilson (eds.). The Sauropods: Evolution and Paleobiology. University of
California Press, Berkeley. 15-49.
Amazonsaurus
Amphicoeliidae Cope, 1877
Reference- Cope, 1877. On Amphicoelias, a genus of saurians from
the Dakota epoch of Colorado. Proceedings of the American Philosophical Society.
17, 242-246.
Amphicoelias
Diplodocimorpha Calvo and Salgado, 1995
Definition- (Limaysaurus tessonei + Diplodocus longus)
(modified from Calvo and Salgado, 1995)
Other definitions- (Rebbachisaurus garasbae + Diplodocus longus)
(modified from Taylor and Naish, 2005)
References- Calvo and Salgado, 1995. Rebbachisaurus tessonei sp.
nov. a new Sauropoda from the Albian-Cenomanian of Argentina; New evidence on
the origin of the Diplodocidae. GAIA. 11, 13-33.
Taylor and Naish, 2005. The phylogenetic taxonomy of Diplodocoidea (Dinosauria:
Sauropoda). PaleoBios. 25(2), 1-7.
Rebbachisauridae Bonaparte, 1997
Definition- (Rebbachisaurus garasbae <- Diplodocus longus)
(Wilson, 2005; modified from Upchurch et al., 2004)
References- Bonaparte, 1997. Rayososaurus agrioensis Bonaparte
1995. Ameghiniana. 34, 116.
Upchurch, Barrett and Dodson, 2004. Sauropoda. in Weishampel, Dodson and Osmolska
(eds.). The Dinosauria (2nd edition). University of California Press, Berkeley.
259-322.
Wilson, 2005. Overview of sauropod phylogeny and evolution. in Curry Rogers
and Wilson (eds.). The Sauropods: Evolution and Paleobiology. University of
California Press, Berkeley. 15-49.
Rebbachisauroidea Bonaparte, 1997 vide Apesteguia, 2007
Definition- (Histriasaurus boscarollii + MACN PV N35 + Rebbachisaurus
garasbae + Limaysaurus tessonei) (after Apesteguia, 2007)
Comments- Apesteguia (2007) erected Rebbachisauroidea to contain a node-based
Rebbachisauridae and their basal relatives such as Histriasaurus and
MACN PV N35, but Rebbachisauridae is a stem-based clade which includes these
latter two taxa anyway. This would leave Rebbachisauroidea as a subgroup of
Rebbachisauridae with identical known content. Since the term itself has limited
utility, has not been used since it was erected, and violates the ICZN by being
inside another superfamily (Diplodocoidea) and a family (Rebbachisauridae),
it is rejected here.
References- Bonaparte, 1997. Rayososaurus agrioensis Bonaparte
1995. Ameghiniana. 34, 116.
Apestegu�a, 2007. The sauropod diversity of the La Amarga Formation (Barremian),
Neuqu�n (Argentina). Gondwana Research. 12, 533-546.
Algoasaurus
Histriasaurus
Zapalasaurus
Nopcsaspondylus
Comahuesaurus Carballido,
Salgado, Pol, Canudo and Garrido, 2012
C. windhauseni Carballido, Salgado, Pol, Canudo and Garrido, 2012
Aptian-Albian-Early Cretaceous
Puesto Quiroga Member of the Lohan Cura Formation, Neuquen, Argentina
Holotype- (MOZ-PV 6722) (at least three individuals) ~tenth-twelfth dorsal
neural arch
Paratypes- ..(MOZ-PV 6627) mid caudal vertebra (143 mm)
..(MOZ-PV 6628) mid caudal vertebra (165 mm)
..(MOZ-PV 6629) mid caudal vertebra
..(MOZ-PV 6630) proximal caudal vertebra (105 mm)
..(MOZ-PV 6631) partial proximal caudal vertebra (143 mm)
..(MOZ-PV 6632) mid caudal vertebra
..(MOZ-PV 6633) mid caudal vertebra (160 mm)
..(MOZ-PV 6634) mid caudal vertebra (166 mm)
..(MOZ-PV 6635) proximal caudal vertebra (90 mm)
..(MOZ-PV 6636) proximal caudal vertebra (127 mm)
..(MOZ-PV 6637) proximal caudal centrum (100 mm)
..(MOZ-PV 6638) mid caudal vertebra (168 mm)
..(MOZ-PV 6639) distal caudal vertebra (225 mm)
..(MOZ-PV 6640) proximal caudal centrum (107 mm)
..(MOZ-PV 6641) distal caudal vertebra (200 mm)
..(MOZ-PV 6642) mid caudal vertebra (210 mm)
..(MOZ-PV 6643) caudal vertebra
..(MOZ-PV 6644) caudal vertebra
..(MOZ-PV 6645) dorsal centrum
..(MOZ-PV 6646) mid caudal vertebra (198 mm)
..(MOZ-PV 6647) caudal vertebra
..(MOZ-PV 6648) proximal caudal vertebra (110 mm)
..(MOZ-PV 6649) mid caudal vertebra (187 mm)
..(MOZ-PV 6650) anterior dorsal vertebra (102 mm)
..(MOZ-PV 6651) dorsal centrum
..(MOZ-PV 6652) neural arch
..(MOZ-PV 6653) dorsal centrum (182 mm) or neural arch
..(MOZ-PV 6654) mid caudal vertebra (192 mm)
..(MOZ-PV 6657) mid-psoterior dorsal neural arch
..(MOZ-PV 6658) incomplete ischium
..(MOZ-PV 6659) pubic fragment
..(MOZ-PV 6660) pubic fragment
..(MOZ-PV 6661) fragmentary femur
..(MOZ-PV 6663) pubic fragment
..(MOZ-PV 6664) humeral fragment
..(MOZ-PV 6665) femur
..(MOZ-PV 6666) fragmentary femur
..(MOZ-PV 6667) pubic fragment
..(MOZ-PV 6669) two pubic fragments
..(MOZ-PV 6670) pubic fragment
..(MOZ-PV 6672) humeral fragment
..(MOZ-PV 6673) humeral fragment
..(MOZ-PV 6675) fragmentary ilium
..(MOZ-PV 6676) fragmentary ischium
..(MOZ-PV 6680) fragmentary ischium
..(MOZ-PV 6711) distal caudal vertebra (220 mm)
..(MOZ-PV 6712) humeral fragment
..(MOZ-PV 6713) partial ischium
..(MOZ-PV 6714) humeral fragment
..(MOZ-PV 6716) fragmentary ischium
..(MOZ-PV 6717) sternal plate
..(MOZ-PV 6719) partial ischium
..(MOZ-PV 6721) fragmentary femur
..(MOZ-PV 6723) humeral fragment
..(MOZ-PV 6727) partial fibula
..(MOZ-PV 6728) femur (1.13 m)
..(MOZ-PV 6729) mid caudal vertebra (180 mm)
..(MOZ-PV 6732) femur
..(MOZ-PV 6733) distal caudal vertebra (210 mm)
..(MOZ-PV 6734) distal caudal vertebra
..(MOZ-PV 6738) mid caudal vertebra (215 mm)
..(MOZ-PV 6740) proximal caudal vertebra
..(MOZ-PV 6741) incomplete first caudal vertebra (185 mm)
..(MOZ-PV 6743) pubis (805 mm)
..(MOZ-PV 6745) proximal caudal vertebra (90 mm)
..(MOZ-PV 6747) dorsal centrum
..(MOZ-PV 6748) proximal caudal vertebra
..(MOZ-PV 6751) dorsal centrum or chevron
..(MOZ-PV 6753) mid caudal vertebra (210 mm)
..(MOZ-PV 6755) femur
..(MOZ-PV 6756) partial posterior dorsal vertebra (183 mm)
..(MOZ-PV 6759) mid caudal vertebra (220 mm)
..(MOZ-PV 6760) proximal caudal vertebra (127 mm)
..(MOZ-PV 6761) femur
..(MOZ-PV 6762) humerus (675 mm)
..(MOZ-PV 6763) incomplete coracoid
..(MOZ-PV 6764) proximal tibia
..(MOZ-PV 6766) mid caudal vertebra (190 mm)
..(MOZ-PV 6767) proximal caudal vertebra (120 mm)
..(MOZ-PV 6778) fragmentary femur
Referred- ..(MOZ-PV 6718) tooth (Salgado et al., 2004)
Diagnosis- (after Carballido et al., 2012) anterior dorsal centra with
strong lateral constriction, resulting in a thin ventral keel; anterior dorsal
vertebrae with long prezygapophyses, in anterior view covering around 3/4 of
the transverse processes; anterior dorsal vertebrae with two spinoprezygapophyseal
laminae; anterior dorsal vertebrae with two spinodiapophyseal laminae, an anterior
and a posterior one; anterior median lamina formed by three different laminae,
the anterior and posterior spinoprezygapophyseal laminae and the anterior spinodiapophyseal;
posterior dorsal centra with centroprezygapophyseal lamina medially divided;
posterior dorsal neural arches with three spinopostzygapophyseal laminae; double
contact between posterior spinodiapophyseal lamina and lateral spinopostzygapophyseal
lamina; proximal caudal vertebrae with well-developed prezygodiapophyseal fossa;
caudal vertebrae with short transverse process; robust humerus (robustness index
0.3); ischium with straight shaft; shaft of ischium forming right angle with
acetabulum; ilial peduncle of ischium without constriction or neck.
Comments- The material was collected in 1995-1996 and 2002, and initially
described as Limaysaurus sp. by Salgado et al. (2004). Carballido et
al. (2012) described the rest of the material once it was prepared, found that
the characters used to refer it to Limaysaurus were unknown in other
rebbachisaurids. Therefore, they erected a new genus for the material, and recovered
Comahuesaurus as sister to Khebbashia. Salgado et al. identified MOZ-PV
6741 as a last sacral vertebra, but it was reidentified as the first caudal
by Carballido et al..
References- Salgado, Garrido, Cocca and Cocca, 2004. Lower Cretaceous
rebbachisaurid sauropods from the Cerro Aguada Leon (Lohan Cura Formation),
Neuquen Province, northwestern Patagonia, Argentina. Journal of Vertebrate Palaeontology.
24(4), 903-912.
Carballido, Salgado, Pol, Canudo and Garrido, 2012. A new basal rebbachisaurid
(Sauropoda, Diplodocoidea) from the Early Cretaceous of the Neuquen Basin; evolution
and biogeography of the group. Historical Biology. 24(6), 631-654.
Khebbashia Fanti, Cau, Cantelli, Hassine
and Auditore, 2015
Definition- (Limaysaurus tessonei + Nigersaurus taqueti
+ Rebbachisaurus garasbae) (Fanti, Cau, Cantelli, Hassine and Auditore,
2015)
Reference- Fanti, Cau, Cantelli, Hassine and Auditore, 2015. New information
on Tataouinea hannibalis from the Early Cretaceous of Tunisia and implications
for the tempo and mode of rebbachisaurid sauropod evolution. PLoS ONE. 10(4),
e0123475.
Katepensaurus Ibiricu,
Casal, Martinez, Lamanna, Luna and Salgado, 2013
K. goicoecheai Ibiricu, Casal, Martinez, Lamanna, Luna and Salgado,
2013
Middle Cenomanian-Turonian, Late Cretaceous
Lower Member of Bajo Barreal Formation of the Chubut Group, Chubut, Argentina
Holotype- (UNPSJB-PV 1007) incomplete frontal, two incomplete anterior cervical
vertebrae with fused ribs, incomplete mid cervical vertebra with fused ribs,
two cervical rib fragments, partial anterior dorsal vertebra, partial anterior
to mid dorsal vertebra, incomplete anterior to mid dorsal neural arch, three
incomplete mid to posterior dorsal vertebrae (175 mm), four incomplete proximal
caudal vertebrae, proximal caudal neural arch, neural arch fragment, several
partial dorsal or proximal caudal neural spines, ?astragalar fragment, ?metapodial
fragment, several fragments
Diagnosis- (after Ibiricu et al., 2013) internal lamina divides lateral
pneumatic fossa of mid to posterior dorsal centrum (also in Dinheirosaurus
and Supersaurus); vertical ridges on lateral surface of mid to posterior
dorsal vertebra, overlying neurocentral junction; pair of laminae in mid to
posterior dorsal parapophyseal centrodiapophyseal fossa; mid to posterior dorsal
transverse processes perforated by elliptical fenestrae (that decrease in size
posteriorly through dorsal series and occupy approximately two-thirds of mediolateral
process width); well-defined, rounded fossae on lateral aspect of mid to posterior
dorsal postzygapophyses.
(after Ibiricu et al., 2015) posterior articular surface of mid to posterior
dorsal centrum with ventral portion wider than dorsal portion, making it teardrop-shaped;
anterior dorsal transverse processes have laterodiapophyseal fossae homologous
with more posterior fenestrae.
Comments- Based on the quarry name, the holotype was discovered in 2005.
Ibiricu et al. (2013) referred Katepensaurus to Rebbachisauridae and
provisionally to Limaysaurinae. Ibiricu et al. (2015) described further material
of the holotype and added the taxon to Whitlock's diplodocoid analysis to find
it emerged as a limaysaurine. However, Fanti et al. (2015) also used a version
of that analysis, and independently coding the originally described material,
recovered Katepensaurus as a basal rebbachisaurine. As this latter analysis
had the benefit of including Tataouinea and information from Rebbachisaurus'
redescription, which result is better supported is uncertain without further
work.
References- Ibiricu, Casal, Martinez, Lamanna, Luna and Salgado, 2013.
Katepensaurus goicoecheai gen. et sp. nov., a Late Cretaceous rebbachisaurid
(Sauropoda, Diplodocoidea) from central Patagonia, Argentina. Journal of Vertebrate
Paleontology. 33(6), 1351-1366.
Fanti, Cau, Cantelli, Hassine and Auditore, 2015. New information on Tataouinea
hannibalis from the Early Cretaceous of Tunisia and implications for the
tempo and mode of rebbachisaurid sauropod evolution. PLoS ONE. 10(4), e0123475.
Ibiricu, Casal, Martinez, Lamanna, Luna and Salgado, 2015. New material of Katepensaurus
goicoecheai (Sauropoda: Diplodocoidea) and its significance for the morphology
and evolution of Rebbachisauridae. Ameghiniana. 52(4), 430-446.
Limaysaurinae Whitlock, 2011
Definition- (Limaysaurus tessonei <- Nigersaurus taqueti)
(modified from Whitlock, 2011
Reference- Whitlock, 2011. A phylogenetic analysis of Diplodocoidea (Saurischia:
Sauropoda). Zoological Journal of the Linnean Society. 161(4), 872-915.
Limaysaurus
Cathartesaura
Rayososaurus
Rebbachisaurinae Bonaparte, 1997 vide
Fanti, Cau, Cantelli, Hassine and Auditore, 2015
Definition- (Rebbachisaurus garasbae <- Limaysaurus tessonei)
(Fanti, Cau, Cantelli, Hassine and Auditore, 2015)
= Nigersaurinae Whitlock, 2011
Definition- (Nigersaurus taqueti <- Limaysaurus tessonei) (modified
from Whitlock, 2011
References- Bonaparte, 1997. Rayososaurus agrioensis Bonaparte
1995. Ameghiniana. 34, 116.
Whitlock, 2011. A phylogenetic analysis of Diplodocoidea (Saurischia: Sauropoda).
Zoological Journal of the Linnean Society. 161(4), 872-915.
Fanti, Cau, Cantelli, Hassine and Auditore, 2015. New information on Tataouinea
hannibalis from the Early Cretaceous of Tunisia and implications for the
tempo and mode of rebbachisaurid sauropod evolution. PLoS ONE. 10 (4), e0123475.
Nigersaurus
Demandasaurus Torcida Fern�ndez-Baldor,
Canudo, Huerta, Montero, Pereda Suberbiola and Salgado, 2011
D. darwini Torcida Fern�ndez-Baldor, Canudo, Huerta, Montero,
Pereda Suberbiola and Salgado, 2011
Late Barremian-Early Aptian, Early Cretaceous
Castrillo la Reina Formation, Burgos, Spain
Holotype- (MDS-RVII) (adult, ~10-12 m) premaxillae, dentary, six teeth (~2.6x.6x?,
~1.5x.6x.4, 1.9x.6x.4, 1.6x.6x.4, 1.5x.6x.4, 1.2x.5x? mm), axis (100 mm), mid
cervical vertebra (270 mm), posterior cervical vertebra, five cervical ribs,
two posterior dorsal vertebrae (150, 150 mm), nine dorsal ribs, first caudal
vertebra (145 mm), second caudal vertebra (92 mm), six proximal caudal vertebrae
(98, 100 mm), eleven mid caudal vertebrae (150, 165, 165, 185, 175, 175 mm),
nine chevrons (315, 355, 370, 340, ~135, 205, 150, 131, 130 mm), ischia, femur
(1.08 m)
Diagnosis- (after Torcida Fernandez-Baldor et al., 2011) teeth ornamented
with longitudinal crests on labial and lingual faces, and bear mesial and distal
carinae; posterior cervical vertebrae have infraprezygapophyseal chamber with
forked vertical accessory lamina; posterior cervical vertebrae have rhombic
accessory structure where centroprezygapophyseal, prezygodiapophyseal and spinoprezygapophyseal
laminae are connected, dorsally to prezygapophyses; presacral centroprezygapophyseal
laminae divided; presence in mid dorsals of two large neural arch pneumatic
foramina that pass completely through neural arch anteroposteriorly; presence
of two large, deep pneumatic cavities, divided by accessory laminae, in proximal
caudal transverse processes; proximal caudal anterior centroparapophyseal, posterior
centroparapophyseal and posterior centrodiapophyseal laminae very wide and make
contact posteriorly and ventrally with diapophysis; two parallel laminae running
in anteroposterior direction, an upper one from prezygapophysis to base of centropostzygapophyseal,
and lower one from base of prezygapophysis to dorsal surface of proximal caudal
centra; two parallel crests running anteroposteriorly on lateral faces of mid-distal
caudal centra.
Comments- Found in 2002-2004. Torcida et al. (2011) added Demandasaurus
to Sereno et al.'s rebbachisaurid matrix and found it to be a nigersaurine.
This general relationship has been recovered in subsequent analyses as well,
though the redescription of Rebbachisaurus has led to the subfamily being
renamed Rebbachisaurinae. Most recently, Fanti et al. (2015) found it to be
sister to Rebbachisaurus+Tataouinea using a version of Whitlock's
diplodocoid analysis.
References- Pereda Suberbiola, Torcida, Izquierdo, Huerta, Montero and
P�rez, 2003. First rebbachisaurid dinosaur (Sauropoda, Diplodocoidea)
from the Early Cretaceous of Spain: Palaeobiogeographical implications. Bulletin
de la Soci�t� G�ologique de France. 174, 471-479.
Torcida Fern�ndez-Baldor, Canudo, Huerta, Montero, Pereda Suberbiola
and Salgado, 2011. Demandasaurus darwini, a new rebbachisaurid sauropod
from the Early Cretaceous of the Iberian Peninsula. Acta Palaeontologica Polonica.
56(3), 535-552.
Fanti, Cau, Cantelli, Hassine and Auditore, 2015. New information on Tataouinea
hannibalis from the Early Cretaceous of Tunisia and implications for the
tempo and mode of rebbachisaurid sauropod evolution. PLoS ONE. 10(4), e0123475.
Rebbachisaurus
Tataouinea Fanti, Cau, Hassine
and Contessi, 2013
T. hannibalis Fanti, Cau, Hassine and Contessi, 2013
Early Albian, Early Cretaceous
Oum ed Diab Member of the Ain el Guettar Formation, Tunisia
Holotype- (ONM DT 1-36) (adult) partial synsacrum (~800 mm), partial first
caudal vertebra, partial second caudal vertebra, incomplete third caudal vertebra,
incomplete fourth caudal vertebra, fifth caudal vertebra, incomplete sixth caudal
vertebra (140 mm), incomplete seventh caudal vertebra (130 mm), eighth caudal
vertebra (130 mm), ninth caudal vertebra (190 mm), tenth caudal vertebra (160
mm), eleventh caudal vertebra (230 mm), twelfth caudal vertebra (230 mm), thirteenth
caudal vertebra (240 mm), fourteenth caudal vertebra (210 mm), fifteenth caudal
vertebra (240 mm), sixteenth caudal vertebra (250 mm), incomplete seventeenth
caudal vertebra (220 mm), partial ilia, proximal ischia
Diagnosis- (after Fanti et al., 2013) elliptical foramen in lateral surface
of fourth sacral neural spine penetrating its camerate sector; proximalmost
five caudal vertebrae with large elliptical pleurocoel (modified after Fanti
et al., 2015); 'lateral lamina' in proximal caudal neural spines is 'inverted
Y'-shaped, formed by spinoprezygapophyseal and spinodiapophyseal laminae merging
in ventral third of spine and bordering a triangular fossa; ischium with large
elliptical foramen in medial surface of ilial peduncle.
Comments- The holotype was discovered in 2011, preliminarily named and
described in 2013, then described in depth two years later. While Fanti et al.
(2013) originally described what they thought were the first five caudal vertebrae,
they later (2015) identified the real second caudal that belongs in the middle
of that series, and eleven subsequent caudals that weren't excavated at the
time. Both papers used a version of Whitlock's diplodocoid analysis to recover
Tataouinea in similar positions within Rebbachisauridae, though the proper
subfamily name changed from Nigersaurinae to Rebbachisaurinae based on Rebbachisaurus
itself being restudied and thus reclassified.
References- Fanti, Cau, Hassine and Contessi, 2013. A new sauropod dinosaur
from the Early Cretaceous of Tunisia with extreme avian-like pneumatization.
Nature Communications. 4, 2080.
Fanti, Cau, Cantelli, Hassine and Auditore, 2015. New information on Tataouinea
hannibalis from the Early Cretaceous of Tunisia and implications for the
tempo and mode of rebbachisaurid sauropod evolution. PLoS ONE. 10(4), e0123475.
Flagellicaudata Harris and Dodson, 2004
Definition- (Dicraeosaurus hansemanni + Diplodocus longus)
(Wilson, 2005; modified from Harris and Dodson, 2004)
Reference- Harris and Dodson, 2004. A new diplodocoid sauropod dinosaur
from the Upper Jurassic Morrison Formation of Montana, USA. Acta Palaeontologica
Polonica. 49(2), 197-210.
Wilson, 2005. Overview of sauropod phylogeny and evolution. in Curry Rogers
and Wilson (eds.). The Sauropods: Evolution and Paleobiology. University of
California Press, Berkeley. 15-49.
Dicraeosauridae Huene, 1927
Definition- (Dicraeosaurus hansemanni <- Diplodocus longus)
(Wilson, 2005; modified from Sereno, 1998)
References- Huene, 1927. Short review of the present knowledge of the
Sauropoda. Memoirs of the Queensland Museum. 9(1), 121-126.
Sereno, 1998. A rationale for phylogenetic definitions, with application to
the higher-level taxonomy of Dinosauria. Neues Jahrbuch f�r Geologie und
Pal�ontologie, Abhandlungen. 210(1), 41-83.
Wilson, 2005. Overview of sauropod phylogeny and evolution. in Curry Rogers
and Wilson (eds.). The Sauropods: Evolution and Paleobiology. University of
California Press, Berkeley. 15-49.
undescribed dicraeosaurid (Gallina, Apesteguia, Haluza, Canale and Otero,
2014)
Late Berriasian-Valanginian, Early Cretaceous
Bajada Colorada Formation, Neuquen, Argentina
Material- (MMCH-Pv coll.) incomplete skull, incomplete mandible, vertebrae
Reference- Gallina, Apesteguia, Haluza, Canale and Otero, 2014. The sauropod
fauna of the Bajada Colorada Formation (Berriasian-Valanginian), Neuquen Province,
Argentina. Jornadas Argentinas de Paleontologia de Vertebrados. Ameghiniana.
51(6) suplemento, 11.
Dyslocosaurus
Suuwassea
Amargasaurus
Brachytrachelopan
Dicraeosaurinae Huene, 1927 vide Janensch, 1929
References- Huene, 1927. Short review of the present knowledge of the
Sauropoda. Memoirs of the Queensland Museum. 9(1), 121-126.
Janensch, 1929. Material und Formegehalt der Sauropoden in der Ausbeute der
Tendaguru-Expedition, 1909-1912 [Material and figured content of sauropods in
the yield of the Tendaguru Expedition, 1909-1912]. Palaeontographica. Supplement
VII(1) 2(1), 3-34.
Dicraeosaurus
Diplodocidae Marsh, 1884
Definition- (Diplodocus longus <- Dicraeosaurus hansemanni)
(Wilson, 2005; modified from Sereno, 1998)
= Apatosauridae Huene, 1927
References- Marsh, 1884. Principal characters of American Jurassic dinosaurs.
Part VII. Diplodocidae, a new family of the Sauropoda. American Journal of Science.
27, 161-167.
Huene, 1927. Short review of the present knowledge of the Sauropoda. Memoirs
of the Queensland Museum. 9(1), 121-126.
Sereno, 1998. A rationale for phylogenetic definitions, with application to
the higher-level taxonomy of Dinosauria. Neues Jahrbuch f�r Geologie und
Pal�ontologie, Abhandlungen. 210(1), 41-83.
Wilson, 2005. Overview of sauropod phylogeny and evolution. in Curry Rogers
and Wilson (eds.). The Sauropods: Evolution and Paleobiology. University of
California Press, Berkeley. 15-49.
Amphicoelias
"Probardodiplodocus"
Apatosaurinae Huene, 1927 vide Janensch,
1929
Definition- (Apatosaurus ajax <- Diplodocus longus)
(modified from Taylor and Naish, 2005)
Other definitions- (Apatosaurus louisae <- Diplodocus longus)
(Wilson and Allain, 2015)
= Apatosaurinae sensu Wilson and Allain, 2015
Definition- (Apatosaurus louisae <- Diplodocus longus)
References- Huene, 1927. Short review of the present knowledge of the
Sauropoda. Memoirs of the Queensland Museum. 9(1), 121-126.
Janensch, 1929. Material und Formegehalt der Sauropoden in der Ausbeute der
Tendaguru-Expedition, 1909-1912 [Material and figured content of sauropods in
the yield of the Tendaguru Expedition, 1909-1912]. Palaeontographica. Supplement
VII(1) 2(1), 3-34.
Taylor and Naish, 2005. The phylogenetic taxonomy of Diplodocoidea (Dinosauria:
Sauropoda). PaleoBios. 25(2), 1-7.
Wilson and Allain, 2015. Osteology of Rebbachisaurus garasbae Lavocat,
1954, a diplodocoid (Dinosauria, Sauropoda) from the early Late Cretaceous-aged
Kem Kem beds of southeastern Morocco. Journal of Vertebrate Paleontology. 35(4),
e1000701. DOI:10.1080/02724634.2014.1000701
Atlantosaurus
Apatosaurus
Brontosaurus
Diplodocinae Marsh, 1884 vide Janensch, 1929
Definition- (Diplodocus longus <- Apatosaurus ajax)
(modified from Taylor and Naish, 2005)
Other definitions- (Diplodocus longus <- Apatosaurus louisae)
(Wilson and Allain, 2015)
= Diplodocinae sensu Wilson and Allain, 2015
Definition- (Diplodocus longus <- Apatosaurus louisae)
References- Marsh, 1884. Principal characters of American Jurassic dinosaurs.
Part VII. Diplodocidae, a new family of the Sauropoda. American Journal of Science.
27, 161-167.
Janensch, 1929. Material und Formegehalt der Sauropoden in der Ausbeute der
Tendaguru-Expedition, 1909-1912 [Material and figured content of sauropods in
the yield of the Tendaguru Expedition, 1909-1912]. Palaeontographica. Supplement
VII(1) 2(1), 3-34.
Taylor and Naish, 2005. The phylogenetic taxonomy of Diplodocoidea (Dinosauria:
Sauropoda). PaleoBios. 25(2), 1-7.
Wilson and Allain, 2015. Osteology of Rebbachisaurus garasbae Lavocat,
1954, a diplodocoid (Dinosauria, Sauropoda) from the early Late Cretaceous-aged
Kem Kem beds of southeastern Morocco. Journal of Vertebrate Paleontology. 35(4),
e1000701. DOI:10.1080/02724634.2014.1000701
Diplodocinae indet. (Lapparent and Zbyszewski, 1957)
Tithonian, Late Jurassic
Bombarral Formation, Portugal
Material- (MG 4819; syntype of Megalosaurus pombali) posterior mid-distal caudal centrum
(MG 4821; syntype of Megalosaurus pombali) anterior mid-distal caudal centrum
(MG 4826; syntype of Megalosaurus pombali) posterior mid-distal caudal centrum
Comments- Lapparent and Zbyszewski (1957) referred several vertebrae to their new species of Megalosaurus, M. pombali,
including "a vertebra broken in two but entirely of the same type,
although slightly smaller (Pl. XXV, fig. 86), was recovered at S�o
Greg�rio de Fanadia (Coll. Geol. Serv.)"
However, Mocho et al. (2016) instead found these did not necessarily
belong to one vertebra and referred them to Diplodocinae indet., as
"the general morphology of these vertebrae is indistinguishable from
that of the middle and posterior caudal vertebrae of the diplodocines Diplodocus, Barosaurus and Tornieria."
References- Lapparent and Zbyszewski, 1957. Les dinosauriens du Portugal. M�moires des Services G�ologiques
du Portugal, nouvelle s�rie. 2, 1-63.
, , and ,
2016.
Systematic review of Late Jurassic sauropods from the Museu Geol�gico collections (Lisboa, Portugal). Journal of Iberian Geology. 42:, 227-250.
Tornieria
Supersaurus
Dinheirosaurus
Leinkupal Gallina, Apesteguia,
Haluza and Canale, 2014
L. laticauda Gallina, Apesteguia, Haluza and Canale, 2014
Late Berriasian-Valanginian, Early Cretaceous
Bajada Colorada Formation, Neuquen, Argentina
Holotype- (MMCH-Pv 63-1) ~seventh caudal vertebra (85 mm)
Paratypes- (MMCH-Pv 63-2/3) incomplete ?sixth cervical vertebra (145
mm), incomplete ?eighth cervical vertebra (203 mm)
(MMCH-Pv 63-4) incomplete ?eleventh cervical vertebra (196 mm)
(MMCH-Pv 63-5) partial ?second dorsal vertebra (140 mm)
(MMCH-Pv 63-6) incomplete ~first/second caudal vertebra (60 mm)
(MMCH-Pv 63-7) incomplete ~twelfth caudal vertebra (90 mm)
(MMCH-Pv 63-8) incomplete ~twentieth caudal vertebra (110 mm)
Diagnosis- (after Gallina et al., 2014) proximal caudal transverse process
extremely elongate (about equal or wider to centrum width) with lateroventral
expansions reinforced by robust dorsal and ventral bars; very robust centroprezygapophyseal
lamina in proximal caudal vertebra; paired pneumatic fossae located on base
of postzygapophysis, opposite to articular side, in proximalmost caudal vertebra.
Comments- This taxon was found by Gallina et al. (2014) to be a basal
diplodocine in two versions of Whitlock's diplodocoid matrices, more basal than
Tornieria in the former but sister to it in the latter. Tschopp et al.'s
(2015) specimen-level diplodocoid analysis also recovered it as a diplodocine
outside the Galeamopus+Diplodocus+Barosaurus clade, but
with Tornieria, Dinheirosaurus and Supersaurus more basal.
References- Gallina, Apesteguia, Haluza and Canale, 2014. A diplodocid
sauropod survivor from the Early Cretaceous of South America. PLoS ONE. 9(5),
e97128.
Tschopp, Mateus and Benson, 2015. A specimen-level phylogenetic analysis and
taxonomic revision of Diplodocidae (Dinosauria, Sauropoda). PeerJ. 3:e857.
Galeamopus Tschopp, Mateus and Benson,
2015
G. hayi (Holland, 1924) Tschopp, Mateus and Benson, 2015
= Diplodocus hayi Holland, 1924
?= Galeamopus "shellensis" Tschopp, 2013
?= Galeamopus pabsti Tschopp and Mateus, 2017
Kimmeridgian-Tithonian, Late Jurassic
Morrison Formation, Wyoming, US
Holotype- (HMNS 175; = CM 662; = CMNH 10670) posterior skull, anterior-posterior
cervical vertebrae, cervical ribs, anterior-posterior dorsal vertebrae, sacrum,
proximal-distal caudal vertebrae, chevrons, scapula, coracoid, sternal plate,
humerus, radius, ulna, carpus, metacarpus, manual phalanges, ilium, pubis, ischium,
tibia, fibula, astragalus, metatarsals
Tithonian, Late Jurassic
Brushy Basin Member of the Morrison Formation, Colorado, US (Marsh-Felch
Quarry 1)
Referred- (USNM 2673; = YPM 1922) incomplete skull, mandible (Marsh, 1884)
Kimmeridgian, Late Jurassic
Salt Wash Member of the Morrison Formation, Wyoming, US (Bone Cabin Quarry,
Howe Quarry)
(AMNH 969) skull, mandibles, atlas, axis (Holland, 1906)
(SMA 0011; proposed holotype of Galeamopus "shellensis") (young
adult) incomplete skull, mandibles, teeth, hyoid, proatlas, atlantal intercentrum
(25 mm), atlantal neurapophyses, axis (129 mm), third cervical vertebra (195
mm), fourth cervical vertebra (264 mm), fifth cervical vertebra (320 mm), sixth
cervical vertebra (388 mm), eighth cervical vertebra (410 mm), ninth cervical
vertebra (430 mm), eleventh cervical vertebra (400 mm), twelfth cervical vertebra
(379 mm), fifteenth cervical vertebra (400 mm), cervical ribs, incomplete first
dorsal vertebra, incomplete second dorsal vertebra (355 mm), partial third dorsal
vertebra, fourth dorsal centrum, partial fifth dorsal vertebra (157 mm), partial
sixth dorsal vertebra, incomplete seventh dorsal vertebra, incomplete eighth
dorsal vertebra, incomplete ninth dorsal vertebra, incomplete tenth dorsal vertebra,
several dorsal ribs, partial sacrum, incomplete scapula (1.375 m), coracoid,
several sternal ribs, humeri (870 mm), radius (601 mm), incomplete ulna (603
mm), carpal, metacarpal I (186 mm), phalanx I-1 (53 mm), manual ungual I (202
mm), metacarpal II (218 mm), phalanx II-1 (78 mm), phalanx II-2 (17 mm), metacarpal
III (230 mm), phalanx III-1 (51 mm), metacarpal IV (208 mm), phalanx IV-1 (35
mm), metacarpal V (180 mm), manual claw sheath?, partial ilium (885 mm), incomplete
pubis (835 mm), partial ischium, incomplete femur (1.37 m), tibia (845 mm),
fibula (850 mm), astragalus (250 mm trans), metatarsal I (119 mm), phalanx I-1
(69 mm), pedal ungual I (190 mm), metatarsal II (145 mm), phalanx II-1 (87 mm),
pedal ungual II (157 mm), metatarsal III (158 mm), phalanx III-1 (88 mm), pedal
ungual III (144 mm), metatarsal IV (162 mm), phalanx ?IV-1 (36 mm), metatarsal
V (148 mm) (Klein and Sander, 2008)
Diagnosis- (after Tschopp et al., 2015) distal end of paroccipital process
curved in lateral view; teeth with paired wear facets; well-developed anteromedial
processes on atlantal neurapophyses, which are distinct from the posterior wing;
atlantal neural arch bears small subtriangular, laterally projecting spur at
its base; posterior wing of atlantal neurapophyses remains of subequal width
along most of its length; axial prespinal lamina develops transversely expanded,
knob-like tuberosity at anterior end; interpostzygapophyseal lamina of mid and
posterior cervical neural arches does not project beyond posterior margin of
neural arch.
Comments- The holotype was discovered in 1902 and initially described
as a new species of Diplodocus (Holland, 1924). AMNH 969 was initially
referred to Diplodocus longus (Marsh, 1884), USNM 2673 to Diplodocus
(Holland, 1906), and SMA 0011 to Diplodocinae indet. (Klein and Sander, 2008).
However, all four of these (plus Lourinha specimen ML 418) were found to group
together by Tschopp et al. (2015) outside the Diplodocus+Barosaurus
clade, so were included in their new genus Galeamopus. While SMA 0011
was said to differ from the holotype in several features which could indicate
it is a separate species within the genus, I follow a differing taxonomic philosophy
and would lump all specimens into G. hayi. Tschopp's (2013) thesis fully
describes the specimen and names it Galeamopus "shellensis",
with AMNH 969 as a referred specimen. While names in theses aren't usually listed
in this website, and this one is only because it was used in the supplementary
information of Tschopp et al.. Tschopp and Mateus (2017) officially name SMA 0011 Galeamopus pabsti and instead refer USNM 2673 to it, with AMNH 969 being G. hayi.
References- Marsh, 1884. Principal characters of American Jurassic dinosaurs.
Part VII. On the Diplodocidae, a new family of the Sauropoda. American Journal
of Science (series 3). 27,160-168.
Holland, 1906. The osteology of Diplodocus Marsh. Memoirs of the Carnegie
Museum. 2, 225-264.
Holland, 1924. The skull of Diplodocus. Memoirs of the Carnegie Museum.
9, 378-403.
McIntosh and Berman, 1975. Description of the palate and lower jaw of the sauropod
dinosaur Diplodocus (Reptilia: Saurischia) with remarks on the nature
of the skull of Apatosaurus. Journal of Paleontology. 49, 187-199.
McIntosh, 1990. Species determination in sauropod dinosaurs with tentative suggestions
for their classification. In Carpenter and Currie (eds.). Dinosaur systematics:
Perspectives and approaches. Cambridge University Press. 53-69.
Klein and Sander, 2008. Ontogenetic stages in the long bone histology of sauropod
dinosaurs. Paleobiology. 34, 247-263.
Tschopp, 2013. Evolution of diplodocid sauropod dinosaurs with emphasis on specimens
from Howe Ranch, Wyoming (USA). MS Thesis. Universidade Nova de Lisboa. 455
pp.
Tschopp, Mateus and Benson, 2015. A specimen-level phylogenetic analysis and
taxonomic revision of Diplodocidae (Dinosauria, Sauropoda). PeerJ. 3:e857.
Tschopp and Mateus, 2017. Osteology of Galeamopus pabsti
sp. nov. (Sauropoda: Diplodocidae), with implications for neurocentral
closure timing, and the cervico-dorsal transition in diplodocids.
PeerJ. 5:e3179.
Kaatedocus Tschopp and Mateus,
2012
K. siberi Tschopp and Mateus, 2012
Kimmeridgian, Late Jurassic
Salt Wash Member of the Morrison Formation, Wyoming, US (Howe quarry)
Holotype- (SMA 0004; "HQ 2") (~14 m, subadult) partial skull,
partial mandibles, proatlas, atlas, axis, third cervical vertebra fused to cervical
ribs (113 mm), fourth cervical vertebra fused to cervical ribs (131 mm), fifth
cervical vertebra fused to cervical ribs (165 mm), sixth cervical vertebra fused
to cervical ribs (194 mm), seventh cervical vertebra fused to cervical ribs
(227 mm), eighth cervical vertebra fused to cervical ribs (245 mm), ninth cervical
vertebra fused to cervical ribs (270 mm), tenth cervical vertebra fused to cervical
ribs (273 mm), eleventh cervical vertebra fused to cervical ribs (298 mm), twelfth
cervical vertebra fused to cervical ribs (314 mm), thirteenth cervical vertebra
fused to cervical ribs (322 mm), fourteenth cervical vertebra fused to cervical
ribs (312 mm)
Referred- (AMNH 7530) skull, mandible, eleven cervical vertebrae and
cervical ribs (Brown, 1935)
(SMA D16-3) partial skull (Schmitt et al., 2013)
Diagnosis- (after Tschopp and Mateus, 2012) U-shaped notch separating
frontals anteriorly (also in Galeamopus); squamosals restricted to postorbital
region; rugose tuberosity marks anterodorsal corner of lateral surface of posterior
cervical vertebrae (also in Suuwassea); posterior margin of prezygapophyseal
articular facet of posterior cervical vertebrae bordered posteriorly by conspicuous
transverse sulcus, separating facet from prezygapophyseal process.
Comments- The holotype was discovered in 1990-1991 and initially referred
to Diplodocus (Ayer, 2000) or Barosaurus (Michelis, 2004) before
being described as the new taxon Kaatedocus by Tschopp and Mateus (2012;
which is also a chapter of Tschopp's 2013 thesis). AMNH 7530 was among the Howe
quarry diplodocids recovered in the early 1900s, identified as Barosaurus
by Brown (1935) and never described. SMA D16-3 was identified as Kaatedocus
based on braincase morphology by Schmitt et al. (2013). This was verified and
AMNH 7530 was also referred to the genus by Tschopp et al.'s (2015) specimen-level
diplodocid analysis.
Tschopp and Mateus (2012) modified Whitlock's diplodocoid matrix and found Kaatedocus
to be a basal diplodocine sister to Tornieria+Barosaurus+Diplodocus.
In Tschopp et al.'s (2015) superior analysis, it was found to be a diplodocine
sister to Barosaurus.
References- Brown, 1935. Sinclair dinosaur expedition, 1934. Natural
History. 36, 2-15.
Ayer, 2000. The Howe Ranch Dinosaurs. Sauriermuseum Aathal. 96 pp.
Michelis, 2004. Taphonomie des Howe Quarry’s (Morrison- Formation, Oberer
Jura), Bighorn County, Wyoming, USA. PhD thesis. University of Bonn. 41 pp.
Tschopp and Mateus, 2012. The skull and neck of a new flagellicaudatan sauropod
from the Morrison Formation and its implication for the evolution and ontogeny
of diplodocid dinosaurs. Journal of Systematic Palaeontology. 11(7), 853-888.
Schmitt, Tschopp, Knoll and Sander, 2013. Paleoneuroanatomy and braincase morphology
indicates the presence of at least two diplodocine taxa (Dinosauria: Sauropoda)
at Howe Ranch (Wyoming, USA). Journal of Vertebrate Paleontology, Program and
Abstracts. 23, 206.
Tschopp, 2013. Evolution of diplodocid sauropod dinosaurs with emphasis on specimens
from Howe Ranch, Wyoming (USA). MS Thesis. Universidade Nova de Lisboa. 455
pp.
Tschopp, Mateus and Benson, 2015. A specimen-level phylogenetic analysis and
taxonomic revision of Diplodocidae (Dinosauria, Sauropoda). PeerJ. 3:e857.
Barosaurus
Diplodocus