This phylogeny is the author's opinion based on his interpretation of the literature. It is however, broadly congruent with recent works. As most of these animals lie outside my area on expertise, my ideas about them are dependant more on the literature than my own knowledge. In particular, I'm unable to examine controversial phylogenetic issues myself and must rely on measures such as character/taxon number and recency of publication to reconcile contradictory cladograms. The taxa discussed here are those which have been referred to Theropoda in the past, but are no longer thought to be members of that clade by the author. In addition, animals considered to be carnivorous dinosaurs before the term Theropoda was invented are included. They are arranged phylogenetically below in what is considered their correct placement today. Ex-theropod taxa are indicated in bold. Sometimes a junior synonym of a taxon which was never considered to be theropod was thought to be a theropod at one time, which explains why taxa like Taniwhasaurus are included. The cladogram is not intended to be an extensive listing of non-theropod taxa, which are only included to flesh it out and provide reference for the ex-theropod taxa. A few taxa do not fit in the cladogram and are dealt with immediately below. "Dinosaurus" is a typo not based on any real remains, while Succinodon is based on invertebrate shells, which is also probably the case for Macrodontophion.

Sauropodomorphs which used to be thought of as theropods (including most named before 1950) are dealt with in the main sauropodomorph section. Ex-sauropodomorphs will be added to this cladogram as well.


Format

Alpha
|--Beta
`--Gamma
   |--Delta
   |*-Epsilon
   `--Zeta
      |?-Eta
      `--Theta 

Based roughly on Mike Keesey's old Dinosauricon site. Beta and Gamma are sister taxa within the clade Alpha Delta is within Gamma, but not a member of Zeta. Epsilon, however, is Gamma incertae sedis. Eta is only tentatively placed within Zeta. Nomina nuda are non-italicized and placed in quotation marks, while unpublished taxa (as recognized by the ICZN) are designated the same way, but followed by an asterisk. Nomina dubia are not distinguished from valid taxa, as which taxa are indeterminate is a subjective question. Taxa in parentheses are technically valid according to the ICZN but not used by most theropod workers.


Cladogram


"Dinosaurus"
Macrodontophion Kuphus? putzeri (= Succinodon putzeri) Gnathostomata
|--Chondrichthyes
| |--Elasmobranchii | | |--Hybodontoidea | | | |--Lonchidiidae | | | | `--Lonchidion montsechi (= Priscavolucris montsechi) | | | `--+--Polyacrodontidae | | | `--+--Acrodontidae | | | `--Hybodontidae | | `--Neoselachii
| `--Holocephali `--Osteichthyes |--Actinopterygii `--Sarcopterygii |--Actinistia | |--Sassenidae | |--Laugiidae | `--Coelacanthiformes | |--Coelacanthus | `--Latimeroidea | |--Latimeriidae | `--Mawsoniidae | `--Indocoelacanthus? sp. (= Kota bird) `--Tetrapoda |--Amphibia | |--Gymnophionia | `--Batrachia | |--Caudata | `--Salientia | |--Triadobatrachus | `--+--Notobatrachidae | `--+*?Stremmeia scabra | |--Mesophryne | `--Anura `--Reptiliomorpha |--Lepospondyli `--Amniota |*-Hoplitosuchus raui |--Synapsida | |--+--Ophiacodontidae | | `--Varanopidae | `--+--Caseasauria | `--Eupelycosauria | Sphenacomorpha | |--Edaphosauridae | `--Sphenacodontia | |--Ianthodon | `--+--Haptodus | `--+--Pantelosaurus | `--Sphenacodontoidea | |--Sphenacodontidae | | |--Cutleria | | `--+--Secodontosaurus | | `--+--Cryptovenator | | `--Sphenacodontinae | | |--+--Ctenospondylus | | | `--Sphenacodon | | `--+--Dimetrodon milleri | | `--+--Dimetrodon limbatus | | `--+--Dimetrodon grandis | | `--Bathygnathus borealis | `--Therapsida `--Sauropsida |--Parareptilia | |--Mesosauridae | `--+--Millerettidae | `--Ankyramorpha | |--Lanthanosuchoidea | `--+--Nyctiphrutetus | |--Bolosauridae | `--+--Procolophonoidea | `--+--Nycteroleter | `--Pareiasauria `--Eureptilia |--Coelostegus `--+--+--Thuringothyris | `--Captorhinidae `--+--Brouffia `--Romeriida |--Paleothyris `--+--Hylonomus `--+--Protorothyrididae `--Diapsida |--Araeoscelidia `--Eosuchia |--Weigeltisauridae | |?-Wapitisaurus | |--Coelurosauravus elivensis | |--Coelurosauravus jaekeli | `--Rautiania `--Neodiapsida |--Galesphyrus |--Kenyasaurus |--Lanthanolania |--Tangasauridae |--Thadeosaurus |--Youngina `--+--Claudiosaurus `--Reptilia |--Lepidosauromorpha | |?-Palaeagama | `--+?-Saurosternon | `--Lepidosauriformes | |--Kuehneosauridae | `--Lepidosauria | |--Rhynchocephalia | `--Squamata | |--Dibamidae | `--Bifurcata/Scleroglossa | |--Gekkonomorpha | | Gekkota | | |--+--Diplodactylidae | | | |--Carphodactylidae | | | `--Pygopodidae | | `--+--Eublepharidae | | `--+--Sphaerodactylidae | | `--+--Gekkonidae | | `--Phyllodactylidae | `--Unidentata/Autarchoglossa | |--Scincoidea | | |--Scincidae | | `--+--Xantusiidae | | `--+--Cordylidae | | `--Gerrhosauridae | `--Episquamata | |--Lacertoidea | | Lacertiformes | | |--Teiioidea | | | |--Gymnophthalmidae | | | `--Teiidae | | `--Lacertibaenia | | |--Lacertidae | | `--Amphisbaenia | | |--Rhineuridae | | `--+--Bipedidae | | |--Blanidae | | |--Cadeidae | | `--+--Trogonophidae | | `--Amphisbaenidae | `--Toxicofera | |--Anguimorpha | | Anguiformes | | |--Goannasauria/Varaniformes | | | |--Shinisauridae | | | `--+--Lanthanotidae | | | `--Varanidae | | `--Anguioidea | | |--Xenosauridae | | `--+--Helodermatidae | | `--+--Anniellidae | | `--Anguidae | |--Iguanomorpha | | Iguania | | |--Acrodonta | | | |--Agamidae | | | `--Chamaeleonidae | | `--+--Tropiduridae | | |--Iguanidae | | |--Leiocephalidae | | |--Crotaphytidae | | |--Phrynosomatidae | | |--+--Hoplocercidae | | | `--Polychrotidae | | |--+--Corytophanidae | | | `--Dactyloidae | | `--+--Liolaemidae | | `--+--Opluridae | | `--Leiosauridae | `--Mosasauriformes | Mosasauria | |--+--Aigialosauridae | | `--Mosasauridae | | |--Mosasaurinae | | `--+--Halisauromorpha | | `--Russellosaurina | | |--Russellosaurus | | |--Tethysaurus | | |--Yaguarasaurus | | `--+--Plioplatecarpinae | | `--Tylosaurinae | | |--Hainosaurus | | `--+--Tylosaurus | | `--+--Taniwhasaurus oweni | | |--Taniwhasaurus? ivoensis | | |--Taniwhasaurus antarcticus | | `--Taniwhasaurus mikasaensis (= "Yezosaurus mikasaensis") | `--+--Dolichosauridae | `--+--Adriosaurus | `--+--Haasiophis | |--Pachyrhachis | `--Serpentes | |--Anomalepididae | `--+--+--Leptotyphlodidae | | `--+--Gerrhopilidae | | |--Typhlopidae | | `--Xenotyphlopidae | `--Oculatophidia | Alethinophidia | |--+--Aniliidae | | `--Tropidophiidae | `--+--Xenophiidae | |--Bolyeriidae | |--Boidae | |--+--Anomochilidae | | |--Cylindrophiidae | | `--Uropeltidae | |--+--Xenopeltidae | | `--+--Lococemidae | | `--Pythonidae | `--Macrostomata | |--+--Acrochordidae | | `--Xenodermatidae | `--+--Pareatidae | `--+--Viperidae | `--+--Homalopsidae | |--Lamprophiidae | |--Elapidae | `--Colubridae `--Archosauromorpha `-----+?-Euryapsida | |?-Ichthyosauromorpha | | |--Hupehsuchia | | `--Ichthyosauriformes | | |--Cartorhynchus | | `--+--Chaohusaurus | | `--Ichthyopterygia | | |--Grippidia | | `--Ichthyosauria | | Longipinnati | | |*-Rachitrema pellati | | |--Cymbospondylidae | | `--Hueneosauria | | |--Mixosauridae | | `--Merriamosauria | | |--Shastasauria | | `--Euichthyosauria | | |--Californosaurus | | `--+--Callawayia | | `--+--Toretocnemidae
| | `--Parvipelvia | | |--Macgowania | | `--+--Hudsonelpidia | | `--Neoichthyosauria | `--+--Thalattosauriformes | |--Eusaurosphargis | |--Hanosaurus | |--Helveticosaurus | `--+--Saurosphargidae | `--Sauropterygia | |--Placodontiformes | `--Eosauropterygia | |--Pachypleurosauria | `--Eusauropterygia | |--Corosauridae | |--Cymatosauria | |--Nothosauroidea | `--+--Yunguisaurus | `--Pistosauria | |--Pistosaurus | `--+--Bobosaurus | `--+--Augustasaurus | `--Plesiosauria | |--Eoplesiosaurus | |--Stratesaurus | |--Rhomaleosauridae | |--Plesiosauroidea | `--Pliosauroidea | Pliosauridae | |--Thalassiodracon | `--+--Hauffiosaurus | `--+--Attenborosaurus | `--+--Marmornectes | `--Thalassophonea | |--Peloneustes | `--+--Simolestes | `--+--Liopleurodon | `--+--Brachaucheninae | |*-Gallardosaurus | `--+*-Pliosaurus archiaci | |*?Pliosaurus? frearsi | |*?Pliosaurus? gamma | |*?Pliosaurus? grandis | |*?Pliosaurus? nitidus | |*-Pliosaurus patagonicus | |*?Pliosaurus? planus | |*-Pliosaurus portentificus | |*?Pliosaurus? simplex | |*?Pliosaurus? suprajurensis | |*-Pliosaurus wosinskii | |*-Pliosaurus? sp. (= "Tapinosaurus") | |--+--Pliosaurus brachydeirus | | `?-Pliosaurus rigauxi (= Cetiosaurus rigauxi) | |--Pliosaurus brachyspondylus | |?-Pliosaurus? irgisensis | |--Pliosaurus macromerus | |--Pliosaurus rossicus | |--Pliosaurus westburyensis | `--+--Pliosaurus kevani | `--+--Pliosaurus carpenteri | `--Pliosaurus funkei |?-Choristodera | |*-Actiosaurus gaudryi | |*-Pachystropheus | |--Lazarussuchus | `--+--Cteniogenys | `--+*-Patricosaurus merocratus | |--Hyphalosauridae | | |--Khurendukhosaurus | | `--+--Shokawa | | `--Hyphalosaurus | |--Monjurosuchus | |--Philydrosaurus | `--Neochoristodera | |--Champsosaurus | `--+--Tchoiria | `--+--Ikechosaurus | `--Simoedosaurus |?-Pantestudines | |?-Eunotosaurus | `-+--Pappochelys | `--+--Odontochelys | `--Testudinata | |--Proganochelys | `--+--Australochelys | `--+--Kayentachelys | `--+--Chengyuchelyidae | `--+--Meiolaniidae | `--Testudines | |--Pleurodira | |--Paracryptodira | `--Cryptodira | |--Trionychia | | |--Carettocheylidae | | `--Trionychidae | | |--+--Cyclanorbinae | | | `--Plastomeninae | | `--Trionychinae | | |--Aspideretes | | `--+--Trionyx | | `--+--Aspideretoides | | `--+--Axestemys splendida | | `--+--Axestemys montinsana | | `--+--Axestemys cerevisia | | `--+--Axestemys byssinus | | `--Axestemys quinni | `--Durocryptodira | |--Testudinoidea | `--Americhelydia | |--Chelydroidea | `--Chelonioidea | |--Cheloniidae | `--Dermochelyoidea | |--Dermochelyidae | `?-Protostegidae | |--Archelon | |--Atlantochelys | |--Pneumatoarthrus peloreus | `--Protostega |*?Simiosauria | |*-Protoavis texensis (in part) | |--Vallesaurus | |--Hypuronector
| `--Drepanosauridae | |--Drepanosaurus | |--Dolabrosaurus | `--+--Megalancosaurus preonensis | `--Megalancosaurus endennae |*?Longisquama insignis `--Thecodontia
|--Protorosauria | `--Protorosaurus `--Parathecodontia |--Tanystropheidae | |--Macrocnemus
| `--+--Langobardisaurus | `--+--+--Cosesaurus aviceps | | `--Sharovipteryx | `--+--Gwyneddosaurus erici | `--+--Amotosaurus
| |--Tanystropheus conspicuus | |--Tanystropheus antiquus | |--Tanystropheus longobardicus (= Tribelesodon longobardicus) | |--Tanystropheus haasi | |--Tanystropheus biharicus | `--Tanystropheus? fossai `--+--Rhynchosauria | |--+--Mesosuchus | | `--Otischalkia | `--+--Howesia | `--+--Eohyosaurus | `--+--Rhynchosaurus | `--Hyperodapedontidae | |--Stenaulorhynchus | |--Mesodapedon | `--+--Bentonyx | `--Hyperodapedontinae | |--Fodonyx | `--+--Ammorhynchus | `--+--Isalorhynchus | `--+--Teyumbaita | `--+*-Hyperodapedon fischeri (Scaphonyx fischeri) | |--Hyperodapedon huenei | `--+--+--Hyperodapedon mariensis | | `--Hyperodapedon sanjuanensis | `--+--Hyperodapedon huxleyi | `--+--Hyperodapedon gordoni | `--+--Hyperodapedon stockleyi | `--Hyperodapedon tikiensis `--+--Allokotosauria | |--Pamelaria | `--+--Azendohsauridae | | |--Azendohsaurus laaroussii | | `--Azendohsaurus madagaskarensis | `--Trilophosauridae | |--Teraterpeton | `--+--Spinosuchus caseanus | `--Trilophosaurus `--+--Prolacerta `--Archosauriformes |*-Ankistrodon indicus |*-Cladeiodon lloydii |*?Energosuchus |*-Jaikosuchus |*?"Teleocrater rhadinus" |*-Teratosaurus? bengalensis |*-Tsylmosuchus jakovleri |*-Tsylmosuchus? donensis |*-Tsylmosuchus? samariensis |*-Vjushkovisaurus |*?Zanclodon laevis |*-Zanclodon? crenatus |*-Zanclodon? schuetzii |*?Zanclodon? silesiacus |--Proterosuchia | |--Proterosuchus | `--"Chasmatosaurus" yuani `--+--Fugusuchus |--Sarmatosuchus `--+--Koilamasuchus `--+--Erythrosuchia |*-Wangisuchus `--+--Euparkeriidae | |--Euparkeria | `--+--Halazhaisuchus | `--Osmolskina `--+--Dorosuchus |*-Dongusuchus |*-Scythosuchus |*-Youngosuchus `--+*-Avalonianus sanfordi |*-Basutodon ferox |*-"Cinizasaurus hunti" |*?"Likhoelesaurus ingens" |*-"Massospondylus" hislopi |*-"Megalosaurus" cloacinus |*-"Megalosaurus" obtusus |*-Paleosaurus cylindrodon |*-Paleosaurus? subcylindrodon |*-Picrodon herveyi |*-"Thecodontosaurus" elizae |*-Yarasuchus |*?Zanclodon? bavaricus |*-Zatomus sarcophagus |--Proterochampsia | Proterochampsidae | |--Proterochampsa | `--+--Cerritosaurus | `--+--Tropidosuchus | `--Rhadinosuchinae | |--Gualosuchus | |--Pseudochampsa | `--+--Chanaresuchus | `--Rhadinosuchus `--+--Doswelliidae | |--Archeopelta | |--Doswellia | |--Tarjadia | `--Vancleavea `--Crurotarsi |*-Erpetosuchidae | |--Erpetosuchus | `--Parringtonia |--Parasuchia | |--"Zanclodon" arenaceus | `--Phytosauridae | |*-Belodon plieningeri | |*?Centemodon sulcatus | |*-Clepsysaurus? fraserianus | |*-Clepsysaurus pennsylvanicus | |*-Clepsysaurus? veatleianus | |*-"Megalosaurus" terquemi | |*-Palaeoctanus appalachianus | |*-Palaeoctanus? aulacodus | |*-Palaeoctanus? dumblianus | |*-Palaeoctanus? orthodon | |*-"Paleosaurus" platyodon | |*-"Paleosaurus" stricklandi | |*-Phytosaurus cylindricodon | |*-Rileyasuchus bristolensis | |*-Rileyasuchus? stutchburyi | |*-Rutiodon? leaii | |*-Suchoprion cyphodon | |*-Suchoprion? sulcidens | |--Wannia | `--+--Parasuchus | `--+--Ebrachosuchus | `--Mystriosuchinae | |--Angistorhinus | |--Brachysuchus | `--+--Proteme | |--Rutiodon (= Palaeonornis struthionoides) | `--Leptosuchomorpha | |--Leptosuchus | `--+--Smilosuchus | `--+--Pravusuchus | `--Mystriosuchini | |--Nicrosaurus | `--+--Mystriosuchus | `--+--Machaeroprosopus | `--Redondasaurus `--Archosauria |*-"Cryptoraptor lockleyi" |*-Jushatyria |*-Lukousaurus yini |*-"Plateosaurus" ornatus |*?Sinosaurus? "shawanensis" |*-Tanystrosuchus posthumus |*-Trialestes romeri |--Pseudosuchia | Dromaeosuchia | |--Ornithosuchia | | Ornithosuchidae | | |--Ornithosuchus longidens | | `--+--Riojasuchus tenuisceps | | `--Venaticosuchus rusconii | `--Suchia | |*?Spondylosoma absconditum | |--Aetosauria | | |*-Aetobarbakinoides | | |--+--Revueltosaurus callenderi | | | `--Revueltosaurus hunti | | `--+?-Acaenasuchus | | |--Aetosauroides | | `--Stagonolepididae | | |--Desmatosuchia | | | |--Stagonolepidinae | | | | |--Polesinesuchus | | | | `--Stagonolepis | | | `--Desmatosuchinae | | | |--"Stagonolepis" olenkae | | | `--+--Neoaetosauroides | | | `--+--+--Calyptosuchus | | | | `--+--Adamanasuchus | | | | `--Scutarx | | | `--Desmatosuchini | | | |--Gorgetosuchus | | | `--+--Longosuchus | | | `--+--Sierritasuchus | | | `--+--Lucasuchus | | | `--Desmatosuchus | | `--Aetosaurinae | | |--Stenomyti | | `--+--Aetosaurus | | |--Coahomasuchus | | `--Typothoracinae | | |--Apachesuchus | | `--+--+--Redondasuchus | | | `--+--Chilenosuchus | | | `--Typothorax | | `--Paratypothoracini | | |--Rioarribasuchus | | `--+--Tecovasuchus | | `--Paratypothorax | `--+--Gracilisuchidae | | |--Gracilisuchus stipinicorum | | `--+--Turfanosuchus | | `--Yonghesuchus | `--+--Nundasuchus | `--+--Ticinosuchus | `--Paracrocodylomorpha | |*-Arganasuchus | |*-Vytshegdosuchus | |--Poposauroidea | | |--Qianosuchus | | `--+*-Arctosaurus osborni | | |*-Bromsgroveia | | |*-Ctenosauriscus | | |*-Hypselorhachis mirabilis | | |--+--Arizonasaurus babbitti | | | `--Xilousuchus | | `--+--Poposauridae | | | |--Poposaurus gracilis | | | `--Poposaurus langstoni | | `--+--Lotosaurus | | `--Shuvosauridae | | |--Sillosuchus | | `--+--Shuvosaurus inexpectatus | | `--Effigia | `--Paracrocodyliformes | |*-Heptasuchus | |--Stagonosuchus | `--+--Prestosuchus | `--+--+--Luperosuchus | | `--Saurosuchus | `--+--Apatosuchus orbitoangulatus | `--+--Batrachotomus | `--+--Fasolasuchus | `--+--Teratosauroidea | | Teratosauridae | | |*-Tikisuchus | | |--Rauisuchus | | `--+--Polonosuchus | | |--Postosuchus | | `--Teratosaurus suevicus | `--Crocodylomorpha | |*-Clarensea gracilis | |*-Pedeticosaurus | |*-Pseudohesperosuchus | |*-Saltoposuchus | |--Dromicosuchus | |--Hesperosuchus | `--+--Sphenosuchus | `--+--Dibothrosuchus | |--Terrestrisuchus | `--+--Litargosuchus | `--+--+--Kayentasuchus | | `?-+--Hallopus victor | | `--Macelognathus vagans | `--+--Jungarrsuchus | `--Crocodyliformes | |*-Kemkemia auditorei | |*-"Maroccanoraptor elbegiensis"* | |--Protosuchia | | Protosuchidae | | |--Hemiprotosuchus | | `--Protosuchus | `--Mesoeucrocodylia | |*-Lisboasaurus estesi | |*-Lusitanisuchus mitracostatus | |*-Razanandrongobe sakalavae | |*-Sinocoelurus fragilis | |--Orthosuchus | `--+--Gobiosuchidae | | |--Gobiosuchus | | `--Zaraasuchus | `--+--Edentosuchus | `--+--+--Zosuchus | | `--+--Sichuanosuchus | | `--+--Shantungosuchus | | `--Neuquensuchus | `--+--Hsisosuchus | `--+--Shartegosuchidae | | |*-Kyasuchus | | |--Nominosuchus | | `--+--Fruitachampsa | | `--+--Adzosuchus | | `--Shartegosuchus | `--Metasuchia | |*-Wanosuchus atresus | |*-Trematochampsidae | | |--Itasuchus | | |--Miadanasuchus | | `--Trematochampsa | |--Notosuchia | | Notosuchiformes | | |*-Brasileosaurus pachecoi | | |--Peirosauriformes | | | |--Uruguaysuchidae | | | | |--Araripesuchus | | | | `--Uruguaysuchus | | | `--+?-Cunampaia simplex | | | |--Stolokrosuchus | | | `--+--"Mahajangasuchidae" | | | | |--Kaprosuchus | | | | `--Mahajangasuchus | | | `--+--Hamadasuchus | | | `--Peirosauridae | | | |--Gasparinisuchus | | | |--Lomasuchus | | | |--Montealtosuchus | | | |--Peirosaurus | | | `--Uberabasuchus | | `--+--Candidodon | | `--Ziphosuchia | | |--Libycosuchus | | `--+--+?-Anatosuchus | | | `--Simosuchus | | `--+--+--Malawisuchus | | | `--Pakasuchus | | `--Gondwanasuchia | | |--Notosuchidae | | | |*-Coringasuchus | | | |--Morrinhosuchus | | | `--+--Notosuchus | | | `--+--Labidiosuchus | | | |--Marilliasuchus | | | `--Sphagesauridae | | | |--+--Adamantinasuchus | | | | `--Yacarerani | | | `--+--Caipirasuchus | | | `--+--Sphagesaurus | | | `--+--Caryonosuchus | | | `--Armadillosuchus | | `--+--Chimaerasuchus | | `--+--Comahuesuchus | | `--Sebecosuchia | | |--Pehuenchesuchus | | `--+--+--Doratodon carcharidens | | | `--Doratodon ibericus | | `--Sebecoidea | | |*-Ilchunaia parca | | |--+--Bergisuchus dietrichbergi | | | |--+--Iberosuchus macrodon | | | | `--Iberosuchus crassiproratus | | | `--Sebecidae | | | |--Lorosuchus nodosus | | | `--+--Barinasuchus arveloi | | | |--Eremosuchus elkoholicus | | | `--+--+--Ayllusuchus fernandezi | | | | `--Bretesuchus bonapartei | | | `--+--Langstonia huilensis | | | |--Sebecus icaeorhinus | | | `--Zulmasuchus querejazus | | `--Baurusuchidae | | |--Cynodontosuchus rothi | | `--+--Pabwehshi pakistanensis (?= Vitakridrinda sulaimani) | | `--+--Pissarrachampsinae | | | |--Pissarrachampsa sera | | | `--Wargosuchus australis | | `--Baurusuchinae | | |--Stratiotosuchus maxhechti | | `--+--Campinasuchus dinizi | | `--+--Baurusuchus albertoi | | `--+--Baurusuchus pachecoi | | `--Baurusuchus salgadoensis | `--Neosuchia | |--+--+--Siamosuchus | | | `--Tethysuchia | | | |--Pholidosauridae | | | | Pholidosaurus | | | `--+--Elosuchidae | | | | |--+--Meridiosuchus | | | | | `--Elosuchus | | | | `--+--Sarcosuchus | | | | `--Terminonaris | | | `--Dyrosauridae | | | |--Chenanisuchus | | | `--+--Sokotosuchus | | | `--+--Phosphatosaurus | | | `--+--Cerrejonisuchus | | | `--+--Arambourgisuchus | | | `--+--Dyrosaurus | | | `--+--Congosaurus | | | |--Hyposaurus | | | `--+--Rhabdognathus | | | `--+--Atlantosuchus | | | `--Guarinisuchus | | `--Thalattosuchia | | |*-Cystosaurus | | |*-"Megalosaurus" mersensis | | |--Teleosauroidea | | | `--Teleosaurus | | `--+--Peipehsuchus | | `--+--Steneosaurus | | `--+--Pelagosaurus | | `--Metriorhynchoidea | | |--Teleidosaurus | | `--+--Eoneustes | | `--Metriorhynchidae | | |--Metriorhynchinae | | | |--Metriorhynchus | | | `--+--Gracilineustes | | | `--+--Maledictosuchus | | | `--+--Rhacheosaurus gracilis (= Megalosaurus gracilis) | | | `--+--Cricosaurus | | | `--Neustosaurus | | `--Geosaurinae | | |--Suchodus | | `--+--Purranisaurus | | `--+--Neptunidraco | | `--+--Tyrannoneustes lythrodectikos | | `--+--Torvoneustes carpenteri | | `--+--+--Geosaurus lapparenti | | | `--+--Geosaurus giganteus | | | `--Geosaurus grandis | | `--+*-Dakosaurus nicaeensis (= Aggiosaurus nicaeensis) | | |--Dakosaurus? manselii | | `--+--Dakosaurus maximus (= Megalosaurus schnaitheimii) | | |--Dakosaurus lissocephalus | | `--Dakosaurus andiniensis | `--+*-"Dakosaurus" amazonicus (= Dinosuchus terror) | |*-Stomatosuchidae | | |--Aegyptosuchus | | |--Laganosuchus | | `--Stomatosuchus | |--Susisuchidae | | |--Susisuchus | | `--Isisfordia | `--+--Paluxysuchus | `--+--Goniopholididae | | |--+--Vectisuchus | | | |--Calsoyasuchus | | | `--Sunosuchus | | `--+--"Goniopholis" baryglyphaeus | | `--+--Goniopholis simus | | `--+--"Goniopholis" stovalli | | |--Eutretauranosuchus | | `--Amphicotylus | `--+*-Bernissartia | Eusuchia | |*-Glichristosuchus | |--+--+--Atoposauridae | | | | |--Theriosuchus | | | | `--+--Alligatorium | | | | |--Alligatorellus | | | | |--Atoposaurus | | | | `--Montsecosuchus | | | `--Paralligatoridae | | | |--Wannchampsus | | | `--+--Shamosuchus | | | `--+--Batrachomimus | | | `--+--Rugosuchus | | | `--Paralligator | | `--+?-Pachycheilosuchus | | |--Allodaposuchidae | | | |--Allodaposuchus | | | |--Arenysuchus | | | `--Lohuecosuchus | | `--Hylaeochampsidae | | |--Acynodon | | `--+--Hylaeochampsa | | `--Iharkutosuchus | |--Borealosuchus | `--Crocodylia | |--Alligatoroidea | `--+--Gavialoidea | `--Crocodyloidea `--Avemetatarsalia Avifilopluma Ornithodira |*?Avipes dillstedtianus |?-Pterosauromorpha | |--Protoavis texensis (in part) | |--Scleromochlus taylori | `--Pterosauria | |--+--Preondactylus | | `--+--Peteinosaurus | | `--+--Raeticodactylus | | `--Eudimorphodontidae | `--Macronychoptera | |--Dimorphodontidae | `--Novialoidea | |--Campylognathoididae | `--Breviquartossa | |*-"Archaeopteryx" "vicensensis" | |*-Laopteryx prisca | |--Rhamphorhynchidae | | `--Rhamphorhynchus muensteri | `--+--Sordes | `--Monofenestrata | |--Wukongopteridae | `--+--Changchengopterus | `--Caelidracones | |*-Eurolimnornis corneti | |--Anurognathidae | `--Pterodactyloidea | |*-Cimoliornis diomedeus | |--+--Germanodactylidae | | `--Euctenochasmatia | | |--Pterodactylidae | | | `--Pterodactylus antiquus | | `--+--"Pterodactylus" longicollum | | `--Ctenochasmatoidea | | |--Gallodactylidae | | `--Ctenochasmatidae | `--+*-Samrukia nessovi | |--Haopterus | `--+--Pteranodontia | | |--Nyctosauridae | | `--+--Alamodactylus | | `--Ornithocheiroidea | | |*-Piksi barbarulna | | |--Pteranodontidae | | `--+--Istiodactylidae | | `--+--Lonchodectidae | | `--+--Aetodactylus | | `--+--Cearadactylus | | `--+--Brasileodactylus | | `--+--Ludodactylus | | `--+--Anhangueridae | | `--Ornithocheiridae | `--Azhdarchoidea | |*-Herbstosaurus pigmaeus | |*-Palaeornis clifti | |--+--Bennettazhia | | `--+--Nemicolopterus | | `--+--"Sinopterus" gui | | `--Tapejaridae | `--+--+--Dsungaripteridae | | `--Thalassodrominae | `--+--Chaoyangopteridae | `--+--Cretornis hlavaci | |--Radiodactylus | `--Azhdarchidae | `?-Palaeocursornis corneti `--Dinosauromorpha |--Lagerpetonidae | |--Lagerpeton chanerensis | `--+--Dromomeron gregorii | `--+--Dromomeron romeri | `--Dromomeron gigas `--Dinosauriformes |*-Lagosuchus talampayensis |--Marasuchus lilloensis `--+*-Agnosphitys cromhallensis |*-Nyasasaurus parringtoni |*-Saltopus elginensis |--Silesauridae | |--Lewisuchus admixtus | `--+--Asilisaurus kongwe | `--+*-Ignotosaurus fragilis | |*-Lutungutali sitwensis | |--Eucoelophysis baldwini | |--Silesaurus opolensis | `--+--Diodorus scytobrachion | |--Sacisaurus agudoensis | `--Technosaurus smalli `-----Avipluma Dinosauria |*-Apatodon mirus |*?Praeornis sharovi |--Ornithischia | |--Pisanosaurus | `--+*?Thecospondylus horneri | |*?Tichosteus? aequifacies | |*?Tichosteus lucasanus | |--Heterodontosauridae | |--Eocursor | `--Genasauria | |--Thyreophora | | |*-Regnosaurus northamptoni | | |--Scutellosaurus | | `--+--Emausaurus | | `--+--Scelidosaurus | | `--Eurypoda | | |--Stegosauria | | | |--"Astrodon" pusillus | | | |--Chialingosaurus kuani | | | `--Craterosaurus pottonensis | | `--Ankylosauria | | |--Sarcolestes leedsi | | `--Struthiosaurus austriacus | `--Neornithischia | |--Lesothosaurus | `--+--Agilisaurus | |*-Fulgurotherium australe | `--+--Hexinlusaurus | |--Leaellynasaurua | |--Yandusaurus | `--+--Kulindadromeus zabaikalicus (= Lepidocheirosaurus natatilis) | `--+--Jeholosauridae | `--+--Othneilosaurus | `--+*-Loncosaurus argentinus | |--Thescelosauridae | `--Cerapoda | |--Ornithopoda | | Clypeodonta | | |--Hypsilophodontidae | | `--Iguanodontia | | |--Gasparinisaura | | `--+--Rhabdodontidae | | |--Tenontosaurus | | `--+--Muttaburrasaurus | | `--Dryomorpha | | |--Dryosauridae | | `--Ankylopollexia | | |--Camptosaurus (?=Symphyrophus musculosus) | | `--Styracosterna | | |*-"Cetiosaurus" brachyurus | | |--Theiophytalia | | `--Hadrosauriformes | | |--Iguanodontidae | | `--Hadrosauroidea | | |--Hypsibema crassicauda | | |--Parrosaurus missouriensis | | `--"Troodon" isfarensis | `--Marginocephalia | |--Pachycephalosauria | `--Ceratopsia | |--+--Chaoyangosauridae | | `--Psittacosauridae | `--Neoceratopasia `--Saurischia |*-Alwalkeria maleriensis (in part) |*-Herrerasauria | Herrerasauridae | |--Staurikosaurus pricei | `--"Herrerasaurinae" | |--Sanjuansaurus gordilloi | `--Herrerasaurus ischigualastensis |*-Chindesaurus bryansmalli |*-Eoraptor lunensis |*-Teyuwasu barbarenai `--Eusaurischia |--Sauropodomorpha `--Theropoda