This phylogeny is the author's opinion based on his interpretation of the literature. It is however, broadly congruent with recent works (e.g. Tykoski, 2005; Carrano and Sampson, 2007; Carrano et al., 2012; Loewen et al., 2013; Brusatte et al., 2014; Foth et al., 2014; Lee et al., 2014; Bell and Chiappe, 2015; Lu et al., 2015). The relationships of modern birds are based on molecular studies.


Format

Alpha
|--Beta
`--Gamma
   |--Delta
   |*-Epsilon
   `--Zeta
      |?-Eta
      `--Theta 

Based roughly on Mike Keesey's old Dinosauricon site. Beta and Gamma are sister taxa within the clade Alpha Delta is within Gamma, but not a member of Zeta. Epsilon, however, is Gamma incertae sedis. Eta is only tentatively placed within Zeta. Nomina nuda are non-italicized and placed in quotation marks, while unpublished taxa (as recognized by the ICZN) are designated the same way, but followed by an asterisk. Nomina dubia are not distinguished from valid taxa, as which taxa are indeterminate is a subjective question. Taxa in parentheses are technically valid according to the ICZN but not used by most theropod workers.


Cladogram


Theropoda
|--Eodromaeus murphi `--+--+?-Dracoraptor hanigani | |--Daemonosaurus chauliodus | `?-Chilesaurus diegosuarezi `--+--Tawa hallae
`--Avepoda
Averostra Eutheropoda |*-"Megalosaurus" woodwardi |*-"Newtonsaurus" cambrensis |*-Velocipes guerichi
|--Coelophysoidea (Podokesauroidea) | |?-Sarcosaurus woodi | |*-Halticosaurus longotarsus | |*-Sarcosaurus? andrewsi | `--+?-Dilophosauridae
| | |?-"Dilophosaurus" sinensis | | |?-Sinosaurus triassicus | | |?-Dracovenator regenti | | |?-Cryolophosaurus ellioti | | `--Dilophosaurus wetherilli | `--+*?"Comanchesaurus kuesi" | |*-Dolichosuchus cristatus | |*-Gojirasaurus quayi (in part) | |*-Lepidus praecisio | |?-Procompsognathus triassicus
| |*-Protoavis texensis (in part) | |--Liliensternus liliensterni | `--+--Lophostropheus airelensis | |?-Zupaysaurus rougieri | `--Coelophysidae (Podokesauridae) | |*-Camposaurus arizonensis | |*?Coelophysis? willistoni | |*-Kayentavenator elysiae | |*-Longosaurus longicollis | |*-Podokesaurus holyokensis | |*-Pterospondylus trielbae | |*-"Syntarsus" "mexicanum" | |--Segisaurinae | | |?-"Syntarsus" kayentakatae | | `--Segisaurus halli
| `--Coelophysinae | |--Panguraptor lufengensis
| `--+--Coelophysis bauri | `?-Coelophysis? rhodesiensis
`--+--Tachiraptor admirabilis `--Neotheropoda
|*-"Allosaurus" medius |*-Allosaurus? trihedrodon |*-Calamospondylus oweni |*-"Capitalsaurus" potens |*-Chienkosaurus ceratosauroides |*-"Coelurosaurus" |*-Diplotomodon horrificus |*-"Elaphrosaurus" iguidiensis |*-"Futabasaurus"
|*-Inosaurus tedreftensis |*-"Katsuyamasaurus" |*?Kelmayisaurus? "gigantus" |*?"Labrosaurus" "huene" |*-"Megalosaurus" dunkeri |*-"Megalosaurus" insignis |*-"Megalosaurus" lonzeensis |*-Megalosaurus? monasterii |*-"Megalosaurus" pombali |*?Megalosaurus? "tibetensis" |*?"Morosaurus" marchei |*-"Ornithocheirus" hilsensis |*-Orthogoniosaurus matleyi |*?Orthogoniosaurus? rawesi |*-Ozraptor subotaii |*-"Prodeinodon" kwangshiensis |*-Prodeinodon mongoliensis |*-"Scrotum humanum" |*-Szechuanosaurus campi |*-Teinurosaurus sauvagei |*-Wakinosaurus satoi |*-Walgettosuchus woodwardi |*?Yangchuanosaurus? "longqiaoensis"
|--Ceratosauria | |*-"Merosaurus newmani" | |*?"Ngexisaurus dapukaensis" | |--Berberosaurus liassicus | |--+?-+--Deltadromeus agilis | | | `--Gualicho shinyae | | |--Spinostropheus gautieri | | `--+--Elaphrosaurus bambergi | | `--Limusaurus inextricabilis
| `--+--Camarillasaurus cirugedae | `--Neoceratosauria
| |--Ceratosauridae
| | |*?Ceratosaurus? roechlingi | | |--Genyodectes serus | | |?-+--Ostafrikasaurus crassiserratus | | | `--Ostafrikasaurus stechowi | | `--+--Ceratosaurus nasicornis | | `--Ceratosaurus meriani | `--Abelisauroidea | |--Eoabelisaurus mefi | `--+--Ligabueino andesi | `--Abelisauria | |*?Betasuchus bredai | |--Noasauridae | | |*-Coeluroides largus | | |*-Compsosuchus solus | | |*-Jubbulpuria tenuis | | |*-Ornithomimoides? barasimlensis | | |*-Ornithomimoides mobilis | | |*-"Sidormimus"* | | |*?Vitakrisaurus saraiki | | |*-Laevisuchus indicus | | |?-Austrocheirus isasii | | `--+--Velocisaurus unicus | | `--+--Masiakasaurus knopfleri | | `--Noasaurus leali | `--+--Kryptops palaios | |*?"Carnosaurus" | |*-Dryptosauroides grandis | |*-Tarascosaurus salluvicus | `--+--Rugops primus | `--Abelisauridae | |--Majungasaurinae | | |--+--Arcovenator escotae | | | `--Genusaurus sisteronis | | `--+--Indosaurus matleyi | | |--Indosuchus raptorius | | |--Majungasaurus crenatissimus | | `--Rajasaurus narmadensis | `--Brachyrostra | |--Xenotarsosaurus bonapartei | `--+--+--Dahalokely tokana | | `--Rahiolisaurus gujaratensis | `--+*-"Bayosaurus pubica" | |--+--Ilokelesia aguadagradensis | | `--+--Ekrixinatosaurus novasi | | `--Skorpiovenator bustingorryi | `--Furileusauria | |--Pycnonemosaurus nevesi | |--Quilmesaurus curriei | |--Viavenator exxoni | `--Carnotaurini | |--Carnotaurinae | | `--Carnotaurus sastrei
| `--Abelisaurinae | |--Abelisaurus comahuensis | `--Aucasaurus garridoi
`--Tetanurae |--Chuandongocoelurus primitivus |?-Monolophosaurus jiangi |*-Allosaurus? tendagurensis |*-Fosterovenator churei |*-Kaijiangosaurus lini |*?Megalosaurus? "dapukaensis" |*-"Megalosaurus" inexpectatus |*-"Megalosaurus" pannoniensis |*?"Mifunesaurus" |*?Phaedrolosaurus ilikensis |*?Prodeinodon? "tibetensis" |*-"Saltriosaurus" `--Orionides |*-Bahariasaurus ingens |*-Cruxicheiros newmanorum |*-Dandakosaurus indicus |*-Ichthyovenator laosensis |*?"Megalosaurus" hungaricus |*-"Metriacanthosaurus" "reynoldsi" |--Megalosauroidea | |--Piatnitzkysauridae | | |--Marshosaurus bicentissimus | | `--+--Condorraptor currumili | | `--Piatnitzkysaurus floresi | `--Megalosauria | |*-Streptospondylus altdorfensis | |--Megalosauridae | | |*?Streptospondylus? cuvieri | | |--Eustreptospondylinae | | | `--Eustreptospondylus oxoniensis | | `--+--Megalosaurinae | | | |?-Duriavenator hesperis | | | `--+--Megalosaurus bucklandii | | | |*-"Megalosaurus" "phillipsi" | | | |*-"Brontoraptor" | | | |*-Edmarka rex | | | `--+--Wiehenvenator albati | | | `--+--Torvosaurus tanneri | | | `--Torvosaurus gurneyi | | `--Afrovenatorinae | | |--Afrovenator abakensis | | |--Dubreuillosaurus valesdunensis | | |--Leshansaurus qianweiensis | | |?-Magnosaurus nethercombensis | | |--Piveteausaurus divesensis | | `?-Poekilopleuron bucklandii | `--Spinosauridae | |--Baryonychinae | | |--Baryonyx walkeri | | |--Suchosaurus cultridens | | |--Suchosaurus? girardi | | `--Cristatusaurus lapparenti | `--Spinosaurinae | |--Irritator challengeri | |--Oxalaia quilombensis | |--Siamosaurus suteethorni | |--Siamosaurus? fusuiensis | `--Spinosaurus aegyptiacus
`--Avetheropoda |*-Datanglong guangxiensis |*-"Tyrannosaurus" "lanpingensis" |*-Unquillosaurus ceibalei
|--Carnosauria | Allosauroidea | |*-Erectopus superbus | |*?Kelmayisaurus petrolicus | |*-Xuanhanosaurus qilixiaensis | |*-"Yuanmouraptor" | |--Metriacanthosauridae | | |--+--Yangchuanosaurus shangyouensis | | | `--Yangchuanosaurus zigongensis | | `--Metriacanthosaurinae | | |--Shidaisaurus jinae | | `--+--Metriacanthosaurus parkeri | | |--Siamotyrannus isanensis | | |--Sinraptor dongi | | `--Sinraptor? hepingensis | `--Allosauria | |--Carcharodontosauridae | | |--Neovenator salerii | | `--+?*"Osteoporosia gigantea"* | | |--+--Altispinax dunkeri | | | |--Concavenator corcovatus | | | |--Eocarcharia dinops | | | `--Sauroniops pachytholus | | `--+--+--Acrocanthosaurus atokensis | | | `--Veterupristisaurus milneri
| | `--+--+?-Labocania anomala | | | `--Shaochilong maortuensis | | `--Carcharodontosaurinae | | |*-"Megalosaurus" chubutensis | | |*?"Megalosaurus" ingens | | |*-Taurovenator violantei | | |--+--Carcharodontosaurus saharicus | | | `--Carcharodontosaurus? iguidensis | | `--Giganotosaurinae | | |--Tyrannotitan chubutensis | | `--+--Mapusaurus roseae | | `--Giganotosaurus carolinii | `--Allosauridae | |--Allosaurus? ferox | |?-Antrodemus valens | |--Creosaurus atrox | |--Epanterias amplexus | |--Labrosaurus lucaris | |--Saurophaganax maximus | `--Allosaurus fragilis
`--Coelurosauria |-Bicentenaria argentina |-Tugulusaurus faciles |?-Gasosaurus constructus |?-Lourinhanosaurus antunesi |?-Sciurumimus albersdoerferi |*-"Beelemodon" |*-Iliosuchus incognitus |*-"Kagasaurus" `--+*-Xinjiangovenator parvus |?-Megaraptora | |*-"Allosaurus" "robustus" | |*-Rapator ornitholestoides | |?-Chilantaisaurus tashuikouensis
| |?-Siats meekerorum
| `--+*-Orkoraptor burkei | |--Fukuiraptor kitadaniensis | `--Megaraptoridae | |--Aerosteon riocoloradensis | |--"Allosaurus" sibiricus | |--Australovenator wintonensis | |--Megaraptor namunhuaiquii | `--Murusraptor barrosaensis `--Tyrannoraptora |*-Aristosuchus pusillus |*-"Arkansaurus fridayi" |*-Bagaraatan ostromi |*-"Dryosaurus" grandis |*?Kakuru kujani
|*?Microvenator? "chagyabi" |*-Mirischia asymmetrica |*-Nedcolbertia justinhofmanni |*-Santanaraptor placidus |*-Sinocalliopteryx gigas |*-"Tonouchisaurus mongoliensis" |*?"Zunityrannus"* |--Tyrannosauroidea (Deinodontoidea) | |*-Timimus hermani | |--Proceratosauridae | | |--Guanlong wucaii | | |--Proceratosaurus bradleyi | | |--Kileskus aristotocus | | `--+--Sinotyrannus kazuoensis | | `--Yutyrannus huali | `--+?-Aviatyrannis jurassica | `--+--+--Calamosaurus foxi | | |--Dilong paradoxus | | `?-Nuthetes destructor | `--+*-Timurlengia euotica | |--+--Stokesosaurus clevelandi | | |--Juratyrant langhami | | `--Eotyrannus lengi | `--+--Alectrosaurus olseni | `--+?-Embasaurus minax | |--Xiongguanlong baimoensis | `--+--+--Dryptosaurus aquilunguis
| | `--Dryptosaurus? macropus | `--+--Raptorex kriegsteini | `--+--Appalachiosaurus montgomeriensis | `--+--Bistahieversor sealeyi | `--Tyrannosauridae (Deinodontidae) | |*-"Alamotyrannus brinkmani" | |*-Aublysodon? lateralis | |*-Deinodon horridus | |*-Deinodon? falculus | |*?Deinodon? grandis | |*-Deinodon? hazenianus | |*-"Ornithomimus" tenuis | |*?Chingkankousaurus fragilis | |--Albertosaurinae | | |--Gorgosaurus libratus | | |--"Albertosaurus" incrassatus | | |--Albertosaurus sarcophagus | | `?-Albertosaurus? periculosus | `--Tyrannosaurinae | |--+--Nanuqsaurus hoglundi | | `--Teratophoneus curriei | `--+--Lythronax argestes | `--+*-Aublysodon mirandus | |--Daspletosaurus torosus | `--+*?Tyrannosaurus? zhuchengensis | |--Zhuchengtyrannus magnus | `--+--+?-Alioramus remotus | | `--Tarbosaurus bataar | `--Tyrannosaurus rex `--+*-Beipiaognathus jii |--Aorun zhaoi |--Zuolong salleei |--Coeluridae | |--Coelurus fragilis | `--Tanycolagreus topwilsoni `--+--Compsognathidae | |--Huaxiagnathus orientalis | `--+--Sinosauropteryx prima | `--Compsognathus longipes |?-Juravenator starki
`--+--Ornitholestes hermanni |--Scipionyx samniticus `-----------Avepectora Maniraptoriformes |*-Aniksosaurus darwini |*-Archaeornithoides deinosauriscus |--Fukuivenator paradoxus |--Arctometatarsalia | Ornithomimosauria | |*?Kinnareemimus khonkaenensis | |*-"Sanchusaurus" | |--Hexing qingyi | `--+--Deinocheirus mirificus | `--+--Pelecanimimus polyodon | `--+--Harpymimidae | | |--Harpymimus okladnikovi | | `--Shenzhousaurus orientalis | `--Ornithomiminae | |*-Valdoraptor oweni | |--Beishanlong grandis | `--+--Garudimimus brevipes | `--Ornithomimidae | |--"Grusimimus tsuru"* | `--+--Sinornithomimus dongi | |--Qiupalong henanensis | |--"Archaeornithomimus" bissektensis | `--+*-"Coelosaurus" antiquus | |*-Rativates evadens | |*-"Saltillomimus rapidus" | |*-"Tototlmimus packardensis" | |--+--Anserimimus planinychus | | `--+--Dromiceiomimus brevitertius | | `--Dromiceiomimus samueli | `--+--Struthiomimus altus | `--+--Gallimimus bullatus | |*-"Orcomimus" | `--+--Ornithomimus velox | `--+--Ornithomimus? sedens | `--+--Archaeornithomimus asiaticus | `--"Gallimimus" "mongoliensis" `--Maniraptora |*-Ilerdopteryx viai |*-Martharaptor greenriverensis Metornithes |*-Bradycneme draculae |*-Kol ghuva |--Alvarezsauroidea | |*?Alnashetri cerropoliciensis | |?-Nqwebasaurus thwazi | `--+?-Haplocheirus sollers | `--Alvarezsauridae | |--+--Achillesaurus manazzonei | | `--Alvarezsaurus calvoi | `--+--Patagonykinae | | |--Patagonykus puertai | | `--Bonapartenykus ultimus | `--+--Linhenykus monodactylus | `--+--Xixianykus zhangi | `--Parvicursorinae | |*-+--Albertonykus borealis | | `--"Ornithomimus" minutus | |--Parvicursor remotus | `--Mononykini | |*-Heptasteornis andrewsi | |--+--Albinykus baatar | | `--Shuvuuia deserti | `--+--Ceratonykus oculatus | `--Mononykus olecranus |--Therizinosauria | |*-"Tiantaiosaurus sifengensis"
| |--Thecocoelurus daviesi
| |--Falcarius utahensis | `--+--Jianchangosaurus yixianensis
| `--Therizinosauroidea
| |*-"Chilantaisaurus" zheziangensis | |--Beipiaosaurus inexpectus
| `--+--Enigmosauridae | | |--Alxasaurus elesitaiensis | | `--Enigmosaurus mongoliensis | `--+--+--Suzhousaurus megatherioides | | `--+--Erliansaurus bellamanus | | `--Neimongosaurus yangi | `--Therizinosauridae | |*-"Nanshiungosaurus" bohlini | |--+--Nothronychus mckinleyi | | `--+-Erlikosaurus andrewsi
| | `--Therizinosaurus cheloniformis | `--+--Nanshiungosaurus brevispinus | `--+--Segnosaurus galbinensis | `--"Nothronychus" graffami `--Pennaraptora |*-Ningyuansaurus wangi |--Oviraptorosauria
| |--+--Incisivosaurus gauthieri | | `--Protarchaeopteryx robusta | `--+--Similicaudipteryx yixianensis | `--+*-Ganzhousaurus nankangensis | |*-Luoyanggia liudianensis | |--Caudipteridae | | `--Caudipteryx zoui | `--+--Avimimidae | | |?-Microvenator celer | | `--Avimimus portentosus | `--Caenagnathoidea | |*-Gigantoraptor erlianensis | |--Caenagnathidae | | |*-Hagryphus giganteus | | |*-Ojoraptorsaurus boerei | | |--Chirostenotes pergracilis | | |--Apatoraptor pennatus | | |--Anzu wyliei | | `--Elmisaurinae | | |--Kuszholia mengi | | |--Elmisaurus rarus | | |--Elmisaurus? elegans | | `--Leptorhynchos gaddisi | `--Oviraptoridae | |*-Shanyangosaurus niupanggouensis | |--Banji long | |--Citipati osmolskae | |--Huanansaurus ganzhouensis | |--Nankangia jiangxiensis | |--Nomingia gobiensis | |--Oviraptor philoceratops | |--Rinchenia mongoliensis | |--Shixinggia oblita | |--Tongtianlong limosus | |--Wulatelong gobiensis | |--Yulong mini | `--"Ingeniinae" | |--Conchoraptor gracilis | |--Heyuannia huangi | |--"Ingenia" yanshini | |--Jiangxisaurus ganzhouensis | |--Khaan mckannai | |--Machairasaurus leptonychus | `--Nemegtomaia barsboldi `--Paraves Eumaniraptora |*?Boreonykus certekorum |*-Cretaaviculus sarysuensis |*-Elopteryx nopcsai |*-Hulsanpes perlei |*-Jinfengopteryx elegans |*-Xiaotingia zhengi |*-Yixianosaurus longimanus |--Deinonychosauria
| Dromaeosauridae (Ornithodesmidae) | |?-Mahakala omnogovae | |*-Dakotaraptor steini | |*?"Kitadanisaurus" | |*-Ornithodesmus cluniculus | |*?Pamparaptor micros | |*-Pyroraptor olympius | |*-Tianyuraptor ostromi | |*-Variraptor mechinorum | |*-Zapsalis abradens | |*-Zhenyuanlong suni | `--+--Unenlagiinae | | |--Austroraptor cabazai | | `--+--Buitreraptor gonzalezorum | | |--Neuquenraptor argentinus | | |?-Rahonavis ostromi | | |--Unenlagia comahuensis | | `--Unenlagia? paynemili | `--+*?Shanag ashile | |--Microraptoria | | |*-"Paleopteryx thomsoni" | | |*?+--Richardoestesia gilmorei | | | `?-+--Richardoestesia? isosceles | | | `--Richardoestesia? asiatica | | |?-Hesperonychus elizabethae | | `--+--Graciliraptor lujiatunensis | | `--+--Changyuraptor yangi | | `--+--Sinornithosaurus millenii | | `--Microraptor zhaoianus | `--+--Bambiraptor feinbergi | `--Eudromaeosauria | |*-"Dromaeosaurus" gracilis | |*-Yurgovuchia doellingi | |--Dromaeosaurinae | | |--Acheroraptor temertyorum | | |--Atrociraptor marshalli | | |--Saurornitholestes langstoni | | |?-Saurornitholestes? sullivani | | |--Tsaagan mangas | | |--Dromaeosaurus albertensis | | `*?Dromaeosauroides bornholmensis | `--Velociraptorinae (Itemirinae) | |--+--"Ichabodcraniosaurus"
| | |--Velociraptor mongoliensis | | |?-Velociraptor? osmolskae | | |--Luanchuanraptor henanensis | | |--Adasaurus mongoliensis | | `--"Koreanosaurus koreanensis" | `--+*-Pneumatoraptor fodori | |--Deinonychus antirrhopus | |--Itemirus medullaris | `--+--Achillobator giganticus | `--Utahraptor ostrommaysorum |--Archaeopterygidae | |?-Anchiornis huxleyi | |--Archaeopteryx lithographica | |?-Aurornis xui | |?-Eosinopteryx brevipenna | `?-Pedopenna daohugouensis |--Troodontidae | |*?+--Paronychodon lacustris | | |--Paronychodon caperatus | | |--Paronychodon asiaticus | | `--Paronychodon portucalensis | |*?Yaverlandia bitholus | |--+--Mei long | | `--Sinovenator changii | `--+--Geminiraptor suarezarum | `--+*-Koparion douglassi | |*-"Saurornithoides" asiamericanus | |--Sinornithoides youngi | `--+--Byronosaurus jaffei | |--Tochisaurus nemegtensis | `--+--+--Xixiasaurus henanensis | | `--Sinusonasus magnodens | `--+--+--Gobivenator mongoliensis | | `--Philovenator curriei | `--+--Urbacodon itemirensis | `--Troodontinae | |*-"Saurornitholestes" robustus | |--+--Borogovia gracilicrus | | `--Saurornithoides mongoliensis | `--+--Zanabazar junior | `--+--Troodon formosus | |--Troodon? bakkeri | `--+--Linhevenator tani | `--Talos sampsoni `--Avialae |?-Scansoriopterygidae | |--Epidexipteryx hui | |--Scansoriopteryx heilmanni | `--Yi qi `--------+*-Dalianraptor cuhe |--Shenzhouraptor sinensis `--Euavialae |*-Bauxitornis mindszentyae |--Balaur bondoc |--Jixiangornis orientalis |--Yandangornis longicaudus |--Zhongjianornis yangi `--Avebrevicauda |--Chongmingia zhengi |--Omnivoropterygiformes | `--Sapeornis chaoyangensis `--------Pygostylia |*-Qiliania graffini |--Confuciusornithiformes | Confuciusornithidae | |*-"Proornis coreae" | |--Changchengornis hengdaoziensis | `--+*-Zhongornis haoae | |*-Confuciusornis chuonzhous | |--Confuciusornis dui | |--Confuciusornis feducciai | |--Confuciusornis zhengi | `--Confuciusornis sanctus `--Ornithothoraces |*-Aberratiodontus wui |*-"Confuciusornis" jianchangensis |*-Hollanda luceria |*-Lectavis bretincola |*?Lenesornis maltshevskyi |*-Platanavis nana |*-Wyleyia valdensis |*-Xinghaiornis lini |--Enantiornithes | |*-Abavornis bonaparti | |*-Catenoleimus anachoretus | |*-Cuspirostrisornis houi | |*-Enantiophoenix electrophyla | |*-"Enantiornis" walkeri | |*?Evgenavis nobilis | |*-Feitianius paradisi | |*-Gracilornis jiufotangensis | |*-Huoshanornis huji | |*-Iberomesornis romerali | |*-"Ichthyornis" minusculus | |*-+--Incolornis silvae | | `--Incolornis martini | |*-Longchengornis sanyanensis | |*-Yuanjiawaornis viriosus | |--Eopengornis martini | |--Explorornis nessovi | |--Noguerornis gonzalezi | |--Pengornis houi | |--Protopteryx fengningensis | `--+*-Alexornis antecedens | |*-Cratoavis cearensis | |*-Grabauornis lingyuanensis | |*-Halimornis thompsoni | |*-Kizylkumavis cretacea | |*-Sazavis prisca | |--Boluochia zhengi | |--Longipteryx chaoyangensis | |--Longipteryx? yangi | |--Shengjingornis yangi | |--Longirostravisidae | | |--Shanweiniao cooperorum | | `--+--Longirostravis hani | | `--Rapaxavis pani | `--+*-Alethoalaornis agitornis | |*-Gurilynia nessovi | |--+--Avisaurus archibaldi | | `--Avisaurus gloriae | |--Cathayornis? aberransis | |--"Cathayornis" chabuensis | |--Cathayornis yandica | |--Concornis lacustris | |--Dalingheornis liweii | |--Dapingfangornis sentisorhinus | |--Dunhuangia cuii | |--Elbretornis bonapartei | |--Elsornis keni | |--Enantiornis leali | |--Eocathayornis walkeri | |--Eoenantiornis buhleri | |--Flexomornis howei | |--Fortunguavis xiaotaizicus | |--Gobipipus reshetovi | |--Gobipteryx minuta | |--Hebeiornis fengningensis | |--Holbotia ponomarenkoi | |--Houornis caudatus | |--Intiornis inexpectatus | |--Jibeinia luanhera | |--Largirostrornis sexdentoris | |--Liaoxiornis delicatus | |--Linyiornis amoena | |--Martinavis cruzyensis | |--Martinavis? minor | |--Martinavis? saltariensis | |--Martinavis? vincei | |--Martinavis? whetstonei | |--Monoenantiornis sihedangia | |--Mystiornis cyrili | |--Nanantius eos | |--Neuquenornis volans | |--Otogornis genghisi | |--Paraprotopteryx gracilis | |--Parvavis chuxiongensis | |--Pterygornis dapingfangensis | |--Sinornis santensis | |--Soroavisaurus australis | |--Xiangornis shenmi | |--Yungavolucris brevipedalis | |--Bohaiornithidae | | |--+--Shenqiornis mengi | | | `--Sulcavis geeorum | | `--+--Zhouornis hani | | `--+--Longusunguis kurochkini | | `--+--Parabohaiornis martini | | `--Bohaiornis guoi | `--Liaoningornithidae | |?-Vorona berivotrensis | |--Eoalulavis hoyasi | `--Liaoningornis longidigitus
`--+*-Archaeornithura meemannae |*-Jiuquanornis niui |*-Parahongshanornis chaoyangensis |*-Tianyuornis cheni |--Archaeorhynchus spathula |--Chaoyangiformes | `--Chaoyangia beishanensis `--Ornithuromorpha |*-Gargantuavis philoinos |*-"Zhyraornis kashkarovi" |--Hongshanornis longicresta |--Jianchangornis microdonta |--Piscivoravis lii |--+--Bellulornis rectusunguis | `--Schizooura lii |--Patagopterygiformes | |--Patagopteryx deferrariisi | `?-Alamitornis minutus `--+--Longicrusavis houi `--+--Songlingornithidae | |--+--Yanornis martini | | `--Yanornis guozhangi | `--+--Yixianornis grabaui | `--Songlingornis linghensis `--+*-Changmaornis houi |*-Guildavis tener |*-Yumenornis huangi |*-+--Zhyraornis kashkarovi | `--Zhyraornis logunovi |--Juehuaornis zhangi |--Iteravis zheni `--+--Gansuiformes | `--Gansus yumenensis `--+*-Cerebavis cenomanica |*-Horezmavis eocretacea |*-Tingmiatornis arctica |--Ichthyornithes | `--Ichthyornis anceps `--+--Hesperornithes | |*-Potamornis skutchi | |--Pasquiaornis hardiei | |--Pasquiaornis? tankei | `--Hesperornithiformes | |--Enaliornis barretti | |--Enaliornis? sedgwicki | |--Enaliornis? seeleyi | `--Hesperornithoidea | |*-Hesperornis? macdonaldi | |*-Judinornis nogontsavensis | |*-Parascaniornis stensioei | |--Baptornithidae | | `--Baptornis advenus | `--Hesperornithidae | |--+?-Brodavis americanus | | |--Brodavis? baileyi | | |--Brodavis? varneri | | `?-Brodavis? mongoliensis | `--+--Fumicollis hoffmani | `--+--Parahesperornis alexi | |?-Hesperornis? mengeli | `--+--Hesperornis bairdi | `--+*?Canadaga arctica | |*-Hesperornis altus | |*-Hesperornis montanus | |--Hesperornis gracilis | |--Hesperornis chowi | |--Hesperornis "lumgairi" | |--Hesperornis regalis | `--+--Hesperornis crassipes | |--Hesperornis rossicus | `--Hesperornis bazhanovi `--Carinatae |--Ambiortiformes | |--Ambiortus dementjevi | |--Apsaravis ukhaana | `--Palintropus retusus `--Aves |*-Apatornis celer |*-Gallornis straeleni |*-Iaceornis marshi |*-Laornis edvardsianus |*-+--Palaeotringa littoralis | `--Palaeotringa vagans |*-"Palaeotringa" vetus
|*-Volgavis marina |--Palaeognathae | |?-Limenavis patagonica | |--Struthionidae | `--Notopalaeognathae | |--Rheidae | `--+--Apterygidae | `--+--Casuariidae | `--+--Dinornithidae | `--Tinamidae `--Neognathae |*-Novacaesareala hungerfordi |*-Qinornis paleocenica |*-Torotix clemensi |--Galloanserae | |--Pangalliformes | | |*-Austinornis lentus | | |--Quercymegapodiidae | | `--Galliformes | | |--Megapodiidae | | `--+--Cracidae | | `--+--Numididae | | `--Phasianidae | `--Anseriformes | |--Anhimidae | `--Anseres | |--Anseranatidae | `--+--Anatalavis rex | `--Anatoidea | |--Vegavis iaai | |--Presbyornithidae | | `--Teviornis gobiensis | `--Anatidae `--Neoaves |--Strisores | |--Caprimulgidae | `--+--+--Steatornithidae | | `--Nyctibiidae | `--+--Podargidae | `--Daedalornithes | |--Aegothelidae | `--Apodiformes | |--Trochillidae | `--Apodi | |--Hemiprocnidae | `--Apodidae `--+--Columbaves | |--Otidimorphae | | |--Musophagidae | | `--+--Cuculidae | | `--Otididae | `--Columbimorphae | |--Columbidae | `--+--Mesitornithidae | `--Pteroclididae `--+--Gruiformes | |--Ralloidea | | |--Rallidae | | `--+--Heliornithidae | | `--Saurothruridae | `--Grues | |--Psophiidae | `--Gruoidea | |--Aramidae | `--Gruidae `--+--Aequorlitornithes | |--+--Mirandornithes | | | |--Podicipedidae | | | `--Phoenicopteridae | | `--Charadriiformes | | |*?Cimolopterygidae | | | |--Cimolopteryx rara | | | |?-"Cimolopteryx" maxima | | | |--+--Lamarqueavis australis | | | | |--Lamarqueavis minima | | | | `--Lamarqueavis petra | | | `--Ceramornis major | | |?-+--Graculavus velox | | | `--Graculavus augustus | | |?-Telmatornis priscus | | |?-"Lonchodytes" pterygius | | |--Charadrii | | | |--+--Charadriidae | | | | `--Haematopoidea | | | | |--Recurvirostridae | | | | `--Haematopodidae | | | `--Chionidi | | | |--Burhinidae | | | `--+--Chionididae | | | `--Pluvianellidae | | `--+--Scolopaci | | | |--Scolopacidae | | | `--Thinocori | | | |--Thinocoroidea | | | | |--Pedionomidae | | | | `--Thinocoridae | | | `--Parrae | | | |--Rostratulidae | | | `--Jacanidae | | `--Lari | | |--Turnicidae | | `--+--Glareolidae | | `--+--Dromadidae | | `--+--Alcidae | | |--Stercorariidae | | `--Larides | | |--Laridae | | |--Sternidae | | `--Rhynchopidae | `--+--Phaethontimorphae | | |--Phaethontiformes | | | |--Lithoptila | | | `--+--Prophaethon | | | `--Phaethontidae | | `--Eurypygoidea | | |--Eurypygidae | | `--Rhynochetidae | `--Natatores | |--Gaviiformes | | |--Neogaeornis wetzeli | | `--Polarornis gregorii | `--+--Procellariimorphae | | |--Sphenisciformes | | `--Procellariiformes | | |?-Lonchodytes estesi | | |--Tytthostonyx glauconiticus | | |--Diomedeidae | | `--+--Oceanitidae | | `--+--Hydrobatidae | | `--Procellariidae | `--Steganograllatores | |--Ciconiidae | `--Pelecaniformes | |--Steganopodes | | |--Fregatidae | | `--Sulida | | |--Sulidae | | `--Phalacrocoracoidea | | |--Anhingidae | | `--Phalacrocoracidae | `--Pelecanimorphae | |--Threskiornithidae | |--Ardeidae | `--Pelecanoidea | |--Pelecanidae | |--Balaenicipitidae | `--Scopidae `--Inopinaves |--Opisthocomidae `--Dendrornithes |--Accipitriformes | |--Cathartidae | `--Accipitres | |--Sagittariidae | `--+--Pandionidae | `--Accipitridae `--Eutelluraves |--+--Strigiformes | | |--Tytonidae | | `--Strigidae | `--Coraciimorphae | |--Coliidae | `--Cavitaves | |--Leptosomidae | `--Eucavitaves | |--Trogonidae | `--Picocoraciae | |--Bucerotiformes | | |--Bucerotidae | | `--Upupides | | |--Phoeniculidae | | `--Upupidae | `--Picodynastornithes | |--Coraciiformes | | |--+--Meropidae | | | `--Coracioidea | | | |--Brachypteraciidae | | | `--Coraciidae | | `--Halcyones | | |--Todidae | | `--Alcedinoidea | | |--Motmotidae | | `--Alcedinidae | `--Piciformes | |--Galbulae | | |--Bucconidae | | `--Galbulidae | `--Pici | |--Capitonoidea | | |--Capitonidae | | `--Rhamphastidae | `--Picoidea | |--Indicatoridae | `--Picidae `--Australavis |--Cariamidae `--Eufalconimorphae |--Falconidae `--Psittacopasserae |--Psittaciformes | |--Nestoridae | `--+--Cacatuidae | `--Psittacidae `--Passeriformes |--Acanthisittidae `--Eupasseres |--Tyranni | |--Eurylaimides | `--Tyrannides | |--Tyrannoidea | `--Furnarioidea `--Passeri |--Menuridae `--+--Climacteridae `--+--Meliphagoidea `--+--Corvoidea `--+--Picathartidae `--Passerida |--Sylvoidea `--+--Muscicapoidea `--Passeroidea