This phylogeny is the author's opinion based on his interpretation of the literature. It is however, broadly congruent with recent works. As most of these animals lie outside my area on expertise, my ideas about them are dependant more on the literature than my own knowledge. In particular, I'm unable to examine controversial phylogenetic issues myself and must rely on measures such as character/taxon number and recency of publication to reconcile contradictory cladograms. The taxa discussed here are those which have been referred to saurischian groups in the past, but are no longer thought to be members of that clade by the author. In addition, animals considered to be carnivorous dinosaurs before the term Theropoda was invented are included, as are taxa considered by an author to be closer to theropods/birds/sauropods/prosauropods than to any other group even if they didn't technically assign the taxon to any of those groups. They are arranged phylogenetically below in what is considered their correct placement today. Ex-saurischian taxa are indicated in bold. Sometimes a junior synonym of a taxon which was never considered to be saurischian was thought to be a saurischian at one time, which explains why taxa like Taniwhasaurus are included. The cladogram is not intended to be an extensive listing of non-saurischian taxa, which are only included to flesh it out and provide reference for the ex-theropod taxa. A few taxa do not fit in the cladogram and are dealt with immediately below. "Dinosaurus" (and perhaps "Aeposaurus") is a typo not based on any real remains, while Succinodon is based on invertebrate shells, which is also probably the case for Macrodontophion.
Sauropodomorphs which used to be thought of as theropods (including
most named before 1950) are dealt with in the main sauropodomorph
section.
Format
Alpha
|--Beta
`--Gamma
|--Delta
|*-Epsilon
`--Zeta
|?-Eta
`--Theta
Based roughly on Mike Keesey's old Dinosauricon site. Beta and Gamma are sister taxa within the clade Alpha Delta is within Gamma, but not a member of Zeta. Epsilon, however, is Gamma incertae sedis. Eta is only tentatively placed within Zeta. Nomina nuda are non-italicized and placed in quotation marks, while unpublished taxa (as recognized by the ICZN) are designated the same way, but followed by an asterisk. Nomina dubia are not distinguished from valid taxa, as which taxa are indeterminate is a subjective question. Taxa in parentheses are technically valid according to the ICZN but not used by most theropod workers.
Cladogram
"Aeposaurus"
"Dinosaurus"
Macrodontophion
Kuphus? putzeri (= Succinodon putzeri)
Wasaibpanchi damani
Gnathostomata
|--Chondrichthyes
| |--Elasmobranchii
| | |--Hybodontiformes
| | | |--Hamiltonichthys
| | | `--+*-Steinbachodontidae
| | | |--Lonchidiidae
| | | | |--Hylaeobatis
| | | | |--Lonchidion
| | | | |--Parvodus
| | | | |--Pristrisodus
| | | | |--Jiaodontus
| | | | |--Diplolonchidion
| | | | `--+--Vectiselachos? montsechi (= Priscavolucris montsechi)
| | | | `--Vectiselachos ornatus
| | | `--+--Lissodus
| | | |--Isanodus
| | | `--Hybodontidae
| | | |--Acrodontinae
| | | `--Hybodontinae
| | `--Neoselachii
| | |--Batoidea
| | `--Selachimorpha
| | |--Squalimorphii
| | `--Galeomorphii
| | |--Heterodontiformes
| | `--+--Orectolobiformes
| | |--Carcharhiniformes
| | `--Lamniformes
| | |--Mitsukurinidae
| | `--+--+--Carcharias
| | | `--Lamnidae
| | `--+--Alopidae
| | `--+--Pseudocarchariidae
| | `--+--Megachasmidae
| | `--+--Cetorhinidae
| | `--Odontaspidae
| `--Holocephali
`--Osteichthyes
|--Actinopterygii
`--Sarcopterygii
|--Actinistia
| |--Sassenidae
| |--Laugiidae
| `--Coelacanthiformes
| |--Coelacanthus
| `--Latimeroidea
| |--Latimeriidae
| `--Mawsoniidae
| `--Indocoelacanthus? sp. (= Kota bird)
`--Tetrapoda
|--Amphibia
| |--Gymnophionia
| `--Batrachia
| |--Caudata
| `--Salientia
`--Pan-Amniota
|--Lepospondyli
`--Amniota
|*-Hoplitosuchus raui
|--Pan-Mammalia
| |--Caseasauria
| `--+--Ophiacodontidae
| `--Eupelycosauria
| Sphenacomorpha
| |--Edaphosauridae
| `--Sphenacodontia
| |--Ianthodon
| `--+--Haptodus
| `--+--Pantelosaurus
| `--Sphenacodontoidea
| |--Sphenacodontidae
| | |--Cutleria
| | `--+--Secodontosaurus
| | `--+--Cryptovenator
| | `--Sphenacodontinae
| | |--+--Ctenospondylus
| | | `--Sphenacodon
| | `--+--Dimetrodon milleri
| | `--+--Dimetrodon limbatus
| | `--+--Dimetrodon grandis
| | `--Dimetrodon borealis
| `--Therapsida
`-----------------------Sauropsida
|--Captorhinidae
`--Romeriida
|--Anthracodromeus longipes
|--Hylonomus lyelli
|--Paleothyris acadiana
|--Protorothyris archeri
`--Diapsida
|--Araeoscelida
| |--Araeoscelis gracilis
| `--Petrolacosaurus kansensis
`--+--Varanopidae
| |--Ascendonanus nestleri
| |--+--Archaeovenator hamiltonensis
| | `--Orovenator mayorum
| `--+--Apsisaurus witteri
| `--+--Mycterosaurinae
| | |--Mycterosaurus longiceps
| | `--+--Elliotsmithia longiceps
| | `--+--Heleosaurus scholtzi
| | `--Mesenosaurus romeri
| `--Varanodontinae
| |--Aerosaurus greenleorum
| `--+--Varanops brevirostris
| `--+--Varanodon agilis
| `--Watongia meieri
`--Neoreptilia
|--Parareptilia
| |--Mesosauridae
| `--+--Millerettidae
| | |--Milleretta rubidgei
| | `--+--Broomia perplexa
| | `--+--Millerosaurus nuffieldi
| | `--+--Millerosaurus ornatus
| | `--Milleropsis pricei
| `--+--Bolosauridae
| | |--Erpetonyx arsenaultorum
| | `--+--Eudibamus cursoris
| | `--+--Belebey vegrandis
| | `--Bolosaurus striatus
| `--+--Australothyris smithi
| `--+--Microleter mckinzieorum
| `--Ankyramorpha
| |--Acleistorhinidae
| | |--Acleistorhinus pteroticus
| | |--Feeserpeton oklahomensis
| | `--+--+--Delorhynchus cifellii
| | | `--Karutia fortunata
| | `--+--Carbonodrcao lundi
| | |--Colobomycter pholeter
| | `--Colobomycter vaughni
| `--+--+--Nyctiphruretidae
| | | |--Abyssomedon williamsi
| | | `--Nyctiphruretus acudens
| | `--Procolophonoidea
| `--Pareiasauromorpha
| |--Pareiasauria
| `--+--Lanthanosuchus watsoni
| `--+--Macroleter poezicus
| `--+--Bashkyroleter bashkyricus
| |--"Bashkyroleter" mesensis
| |--Emeroleter levis
| |--Nycteroleter ineptus
| `--Rhipaeosaurus tricuspidens
`--+?-Simiosauria
| |--Hypuronector limnaios
| `--+--Vallesaurus cenensis
| `--+--Drepanosauridae
| |--Dolabrosaurus aquatilis
| `--+--+--Protoavis texensis (in part)
| | |--Megalancosaurus endennae
| | `--Megalancosaurus preonensis
| |--Skybalonyx skapter
| `--+--Ancistronychus paradoxus
| |--Avicranium renestoi
| `--Drepanosaurus unguicaudatus
|*?Longisquama insignis
`--Eosuchia
|--Weigeltisauridae
| |--Coelurosauravus elivensis
| |--Glaurung schneideri
| `--+--Weigeltisaurus jaekeli
| `--+--Rautiania alexandri
| `--Rautiania minichi
|--Acerodontosaurus piveteaui
|--Claudiosaurus germaini
|--Hovasaurus boulei
|--Saurosternon bainii
|--Thadeosaurus colcanapi
|--Youngina capensis
`--Reptilia
|*-Albisaurus albinus
|*-Stremmeia scabra
|--Palaeagama vielhaueri
|--Pan-Lepidosauria
| |--Taytalura alcoberi
| |--+--Paliguana whitei
| | |--Marmoretta oxoniensis
| | `--Fraxinisaura rozynekae
| `--Lepidosauria
| |--Sophineta cracoviensis
| |--Rhynchocephalia
| `--+--+--Megachirella wachtleri
| `--+--Ardeosaurus brevipes
| |--Bavarisaurus macrodactylus
| |--Hoyalacerta sanzi
| |--Huehuecuetzpalli mixtecus
| |--+--Oculudentavis khaungraae
| | `--Oculudentavis naga
| `--+--Eichstattisaurus schroederi
| |--Scandensia ciervensis
| `--Squamata
| |*-Alamitornis minutus
| |--Dibamia
| | `--Dibamidae
| `--Scleroglossa
| |--Gekkonomorpha
| | |--Tepexisaurus tepexii
| | `--+--Gobekko cretacicus
| | `--Gekkota
| | |--+--Diplodactylidae
| | | `--+--Carphodactylidae
| | | `--Pygopodidae
| | |--Eublepharidae
| | `--+--Sphaerodactylidae
| | `--+--Gekkonidae
| | `--Phyllodactylidae
| `--Autarchoglossa
| |--Scincoidea
| | |--Dorsetisaurus purbeckensis
| | |--Bainguidae
| | | |--Paramacellodus oweni
| | | `--+--Bainguis parva
| | | `--Parmeosaurus scutatus
| | `--+--Pseudosaurillus becklesi
| | |--Yabeinosaurus tenuis
| | `--+--+--Eolacerta robusta
| | | `--Meyasaurus diazromerali
| | `--+--Scincidae
| | `--+--+--Becklesius hoffstetteri
| | | `--+--+--Globaura venusta
| | | | `--+--Eoxanta lavertifrons
| | | | `--Myrmecodaptria microphagos
| | | `--+--Palaeoxantusia kyrentos
| | | `--Xantusiidae
| | `--+--+--Ornatocephalus metzleri
| | | `--Sakurasaurus shokawensis
| | `--+--Cordylidae
| | `--Gerrhosauridae
| `--Episquamata
| |--Lacertiformes
| | |--Teiioidea
| | | |--Chamops segnis
| | | |--Gymnophthalmidae
| | | |--Teiidae
| | | `--Polyglyphanodontidae
| | `--Lacertibaenia
| | |--Lacertidae
| | `--+--Sineoamphisbeana hexatabularis
| | `--Amphisbaenia
| | |--Rhineuridae
| | `--Amphibaenoidea
| | |--Bipedidae
| | `--+--Blanidae
| | |--Cadeidae
| | `--+--Trogonophidae
| | `--Amphisbaenidae
| `--Toxicofera
| |--+--Iguanomorpha
| | | |--+--+--Ctenomastax parva
| | | | | `--Temujinia ellisoni
| | | | `--+--Anchaurosaurus gilmorei
| | | | `--Zapsosaurus sceliphros
| | | `--Iguania
| | | |*-Igua minuta
| | | |--Chamaeleontiformes
| | | | |--Isodontosaurus gracilis
| | | | `--+--Priscagamidae
| | | | | |--Phrynosomimus asper
| | | | | `--+--Mimeosaurus crassus
| | | | | `--Priscagama gobiensis
| | | | `--Acrodonta
| | | | |--Agamidae
| | | | `--Chamaeleonidae
| | | `--Pleurodonta
| | | |--Tropiduridae
| | | |--Iguanidae
| | | |--Leiocephalidae
| | | |--Crotaphytidae
| | | |--Phrynosomatidae
| | | |--Hoplocercidae
| | | |--Polychrotidae
| | | |--Corytophanidae
| | | |--Dactyloidae
| | | `--+--Liolaemidae
| | | `--+--Opluridae
| | | `--Leiosauridae
| | `--Anguimorpha
| | Anguiformes
| | |--+--Monstersauria
| | | | |--Parasaniwa wyomingensis
| | | | `--+--Colpodontosaurus cracens
| | | | `--+--Parviderma inexacta
| | | | `--+--Paraderma bogerta
| | | | |--Primaderma nessovi
| | | | `--+--Eurheloderma gallicum
| | | | |--Gobiderma pulchrum
| | | | `--+--Palaeosaniwa canadensis
| | | | `--+--Estesia mongoliensis
| | | | `--+--Lowesaurus matthewi
| | | | `--Helodermatidae
| | | `--+--Restes rugosus
| | | `--Anguioidea
| | | |--Xenosauridae
| | | `--+--Carusia intermedia
| | | `--+--Diploglossinae
| | | |--Anguidae
| | | `--Anniellidae
| | `--Goannasauria
| | |--Shinisauria
| | | |--Dalinghosaurus longidigitus
| | | `--+--Merkurosaurus ornatus
| | | `--+--Bahndwivici ammoskius
| | | `--Shinisauridae
| | `--+--Eosaniwa koehni
| | `--+--Paravaranus angustifrons
| | |--Varaniformes
| | | |--Proplatynotia longirostrata
| | | `--+--Aiolosaurus oriens
| | | |--Necrosaurus spp.
| | | |--+--Ovoo gurvel
| | | | `--+--Cherminotus longifrons
| | | | `--Lanthanotidae
| | | `--+--+--Saniwides mongoliensis
| | | | `--Telmasaurus grangeri
| | | `--+--Saniwa ensidens
| | | `--Varanidae
| | `--Mosasauriformes
| | |--Aphanizocnemus libanensis
| | |--Carsosaurus marchesetti
| | `--Mosasauria
| | |--Adriosaurus suessi
| | |--Dolichosaurus longicollis
| | |--+--Coniasaurus crassidens
| | | |--Coniasaurus gracilodens
| | | `--Daliasaurus turneri
| | `--Mosasauroidea
| | |--Aigialosaurus dalmaticus
| | |--Judeasaurus tchernovi
| | |--Russellosaurus coheni
| | `--+--Opetiosaurus bucchichi
| | |--+--Eidolosaurus trauthi
| | | `--Pontosaurus? kornhuberi
| | `--+--Pontosaurus lesuerii
| | |--Tethysaurus nopcsai
| | |--Yaguarasaurus columbianus
| | `--Mosasauridae
| | |--Halisaurinae
| | | |--Eonatator sternbergii
| | | |--Halisaurus arambourgi
| | | `--Halisaurus platyspondylus
| | `--Natantia
| | |--Mosasaurinae
| | | |--Kourisodon puntledgensis
| | | `--+--+--Goronyosaurus nigeriensis
| | | | |--Plesiotylosaurus crassidens
| | | | `--+--Prognathodon overtoni
| | | | `--Prognathodon solvayi
| | | `--+--Ectenosaurus clidastoides
| | | `--+--Clidastes liodontus
| | | `--+--Globidens alabamaensis
| | | |--Mosasaurus hoffmanni
| | | `--Plotosaurini
| | | |--Mosasaurus? lemonnieri
| | | `--+--Moanasaurus manahouangae
| | | `--+--Plotosaurus bennisoni
| | | `--Plotosaurus tuckeri
| | `--+--Plioplatecarpinae
| | | `--Plioplatecarpus primeavus
| | `--Tylosaurinae
| | |--Platycarpus tympaniticus
| | `--+--Tylosaurus proriger
| | |--+--Tylosaurus? nepaeolicus
| | | `--Hainosaurus bernardi
| | `--+--Taniwhasaurus oweni
| | |--Taniwhasaurus? ivoensis
| | |--Taniwhasaurus antarcticus
| | `--Taniwhasaurus mikasaensis (= "Yezosaurus mikasaensis")
| `--Ophidia
| |--Eophis underwoodi
| `--+--Diablophis gilmorei
| |--Portugalophis lignites
| `--+--Parviraptor estesi
| `--+--+--Najash rionegrina
| | `--Coniophis precedens
| `--+--+--Dinilysia patagonica
| | `--Tetrapodophis amplectus
| `--Serpentes
| |--+--Leptotyphlodidae
| | `--Typhlopoidea
| | |--Gerrhopilidae
| | |--Typhlopidae
| | `--Xenotyphlopidae
| |--Anomalepididae
| `--Oculatophidia
| |--Madtsoiidae
| | |--Sanajeh indicus
| | `--+--Wonambi naracoortensis
| | `--Yurlunggur camfieldensis
| |--Simoliophiidae
| | |--Haasiophis terrasanctus
| | `--+--Eupodophis descouensis
| | `--Pachyrhachis problematicus
| `--Alethinophidia
| |--Amerophidia
| | |--Aniliidae
| | `--+--Kataria anisodonta
| | `--Tropidophiidae
| `--Afrophidia
| |--Henophidia
| | |--Xenophiidae
| | |--Bolyeriidae
| | |--Boidae
| | |--Uropeltoidea
| | | |--Uropeltidae
| | | `--+--Cylindrophiidae
| | | `--Anomochilidae
| | `--Pythonoidea
| | |--Xenopeltidae
| | `--+--Lococemidae
| | `--Pythonidae
| `--Caenophidia
| |--Acrochordidae
| `--Colubroides
| |--Xenodermatidae
| `--Colubriformes
| |--Pareatidae
| `--Endogylptodonta
| |--Viperidae
| `--+--Homalopsidae
| `--+--Elapoidea
| | |--Cyclocorus lineanus
| | `--+--Micrelaps bicoloratus
| | `--+--+--Buhoma spp.
| | | `--Elapidae
| | `--+--Pseudaspididae
| | `--+--+--Prosymnidae
| | | `--+--Oxyrhabdium leporinum
| | | `--Psammophiidae
| | `--+--Atractaspidiidae
| | `--+--Lamprophiidae
| | `--Pseudoxyrophidae
| `--Colubroidea
| |--+--Dipsadidae
| | `--Pseudoxenodontidae
| `--+--Natricidae
| `--+--Sibynophiidae
| `--+--Colubridae
| `--+--Calamariidae
| `--Grayiidae
|*-Euryapsida
| |?-Ichthyosauromorpha
| | |--Hupehsuchia
| | `--Ichthyosauriformes
| | |--Cartorhynchus
| | `--+--Chaohusaurus
| | `--Ichthyopterygia
| | |--Grippidia
| | `--Ichthyosauria
| | Longipinnati
| | |*-Rachitrema pellati
| | |--Cymbospondylidae
| | `--Hueneosauria
| | |--Mixosauridae
| | `--Merriamosauria
| | |--Shastasauria
| | `--Euichthyosauria
| | |--Californosaurus
| | `--+--Callawayia
| | `--+--Toretocnemidae
| | `--Parvipelvia
| | |--Macgowania
| | `--+--Hudsonelpidia
| | `--Neoichthyosauria
| `--+--Thalattosauriformes
| |--Eusaurosphargis
| |--Hanosaurus
| |--Helveticosaurus
| `--+--Saurosphargidae
| `--Sauropterygia
| |--Placodontiformes
| | |--Palatodonta
| | `--Placodontia
| | |--Paraplacodus
| | |--+--Placodus gigas (Thecodontosaurus latespinatus paralectotype)
| | | `--Placodus inexpectatus
| | `--Cyamodontoidea
| `--Eosauropterygia
| |--Pachypleurosauria
| `--Eusauropterygia
| |--Corosauridae
| |--Cymatosauria
| |--Nothosauroidea
| `--+--Yunguisaurus
| `--Pistosauria
| |--Pistosaurus
| `--+--Bobosaurus
| `--+--Augustasaurus
| `--Plesiosauria
| |--Eoplesiosaurus
| |--Stratesaurus
| |--Rhomaleosauridae
| |--Plesiosauroidea
| `--Pliosauroidea
| Pliosauridae
| |--Thalassiodracon
| `--+--Hauffiosaurus
| `--+--Attenborosaurus
| `--+--Marmornectes
| `--Thalassophonea
| |--Peloneustes
| `--+--Simolestes
| `--+--Liopleurodon
| `--+--Brachaucheninae
| | |--Brachauchenius
| | |--Kronosaurus
| | `--Megacephalosaurus
| `--+--Gallardosaurus
| `--+*-Pliosaurus archiaci
| |*?Pliosaurus? frearsi
| |*?Pliosaurus? gamma
| |*?Pliosaurus? grandis
| |*?Pliosaurus? nitidus
| |*-Pliosaurus patagonicus
| |*?Pliosaurus? planus
| |*-Pliosaurus portentificus
| |*?Pliosaurus? simplex
| |*?Pliosaurus? suprajurensis
| |*-Pliosaurus wosinskii
| |*-Pliosaurus? sp. (= "Tapinosaurus")
| |--+--Pliosaurus brachydeirus
| | `?-Pliosaurus rigauxi (= Cetiosaurus rigauxi)
| |--Pliosaurus brachyspondylus
| |?-Pliosaurus? irgisensis
| |--Pliosaurus macromerus
| |--Pliosaurus rossicus
| |--Pliosaurus westburyensis
| `--+--Pliosaurus kevani
| `--+--Pliosaurus carpenteri
| `--Pliosaurus funkei
|*-Choristodera
| |*-Actiosaurus gaudryi
| |*-Pachystropheus
| |--Lazarussuchus
| `--+--Cteniogenys
| `--+*-Patricosaurus merocratus
| |--Hyphalosauridae
| | |--Khurendukhosaurus
| | `--+--Shokawa
| | `--Hyphalosaurus
| |--Monjurosuchus
| |--Philydrosaurus
| `--Neochoristodera
| |--Champsosaurus
| `--+--Tchoiria
| `--+--Ikechosaurus
| `--Simoedosaurus
`--Pan-Archosauria
|*-Prolacertoides jimusarensis
|*-Pan-Testudines
| |?-Eunotosaurus
| `-+?-Pappochelys
| `--+--Odontochelys
| `--Testudinata
| |--Proganochelys
| `--+--Australochelys
| `--+--Kayentachelys
| `--+--Chengyuchelyidae
| `--+--Meiolaniidae
| `--Testudines
| |--Pleurodira
| |--Paracryptodira
| `--Cryptodira
| |--Trionychia
| | |--Carettocheylidae
| | `--Trionychidae
| | |--+--Cyclanorbinae
| | | `--Plastomeninae
| | `--Trionychinae
| | |--Aspideretes
| | `--+--Trionyx
| | `--+--Aspideretoides
| | `--+--Axestemys splendida (Dakotaraptor steini types in part)
| | `--+--Axestemys montinsana
| | `--+--Axestemys cerevisia
| | `--+--Axestemys byssinus
| | `--Axestemys quinni
| `--Durocryptodira
| |--Testudinoidea
| `--Americhelydia
| |--Chelydroidea
| `--Chelonioidea
| |--Cheloniidae
| `--Dermochelyoidea
| |--Dermochelyidae
| `?-Protostegidae
| |--Archelon
| |--Atlantochelys
| |--Pneumatoarthrus peloreus
| `--Protostega
|--Aenigmostropheus parringtoni
|--Jesairosaurus lehmani
`--Archosauromorpha
|*-"Thecodontosaurus" primus
|*-Zanclodon? "antiquus"
|--Protorosauria
| `--Protorosaurus speneri
`--Parathecodontia
|--Prolacertidae
| |--Kadimakara australiensis
| `--Prolacerta broomi
`--+*?Kuehneosauridae
| |--Icarosaurus siefkeri
| |--Pamelina polonica
| |--Kuehneosaurus latus
| `--Kuehneosuchus latissimus
|--+*-Trachelosaurus fischeri
| |--+--+--Czatkowiella harae
| | | `--+--Ozimek volans
| | | `?-Sharovipteryx mirabilis
| | `--+--Fuyuansaurus acutirostris
| | `--Dinocephalosauridae
| | |--Dinocephalosaurus orientalis
| | |--Pectodens zhenyuensis
| | `--Protanystropheus antiquus
| `--Tanystropheidae
| |*-Elessaurus gondwanoccidens
| |*-Exilisuchus tubercularis
| |*-Macrocnemus? obristi
| |--+--Macrocnemus bassanii
| | `--Macrocnemus fuyuanensis
| `--+--+--Amotosaurus rotfeldensis
| | `--Augustaburiania vatagini
| `--+*-Gwyneddosaurus erici
| |--+--Langobardisaurus pandolfii
| | `?-Cosesaurus aviceps
| `--+*-Raibliania calligarisi
| |--Sclerostropheus fossai
| `--+--Tanystropheus conspicuus
| |--Tanystropheus longobardicus (= Tribelesodon longobardicus)
| |--Tanystropheus hydroides
| |--Tanystropheus haasi
| `--Tanystropheus biharicus
`--Crocopoda
|--Allokotosauria
| |--Azendohsauridae
| | |--Pamelaria dolichotrachela
| | `--+--Shringasaurus indicus
| | `--+--Azendohsaurus laaroussii
| | `--Azendohsaurus madagaskarensis
| `--Trilophosauridae
| |--Teraterpeton hrynewichorum
| `--+--Trilophosaurus buettneri
| |--Trilophosaurus dornorum
| |--Trilophosaurus phasmalophos
| `--Trilophosaurus caseanus (= Spinosuchus caseanus)
`--+--Rhynchosauria
| |--+--Mesosuchus browni
| | |--Noteosuchus colletti
| | `--Otischalkia elderae
| `--+--Howesia browni
| `--+--Eohyosaurus wolvaardi
| `--+--Rhynchosaurus articeps
| `--Hyperodapedontidae
| |--Stenaulorhynchinae
| | |--Ammorhynchus navajoi
| | `--+--Mesodapedon kuttyi
| | `--+--Brasinorhynchus mariantensis
| | `--Stenaulorhynchus stockleyi
| `--+--Langeronyx brodiei
| |--Bentonyx sidensis
| |--Fodonyx spenceri
| `--Hyperodapedontinae
| |--Isalorhynchus genovefae
| `--+--Oryctorhynchus bairdi
| `--+*-Hyperodapedon fischeri (= Scaphonyx fischeri)
| |--Hyperodapedon gordoni
| `--+--Hyperodapedon sanjuanensis
| `--+--Hyperodapedon huxleyi
| `--+--Hyperodapedon mariensis
| `--+--Hyperodapedon tikiensis
| `--+--Hyperodapedon stockleyi
| `--+--Hyperodapedon sulcognathus
| `--Hyperodapedon huenei
`--+--Boreopricea funerea
`--+*-Ankistrodon indicus
|*-Cladeiodon lloydii
|*-Teratosaurus? bengalensis
|*-Zanclodon? crenatus
|*-Zanclodon? schuetzii
|*-"Zanclodon" silesiacus
|--Teyujagua paradoxa
`--+*-Blomosuchus georgii
|--Tasmaniosaurus triassicus
`--Archosauriformes
|*-Chasmatosuchus? vjushkovi
|*-Eorasaurus olseni
|*-+--Chasmatosuchus magnus
| |--Chasmatosuchus rossicus
| |--Tsylmosuchus jakovleri
| `--Tsylmosuchus? donensis
|--Proterosuchia
| |*-Archosaurus rossicus
| |--Proterosuchus fergusi
| `--+--Proterosuchus alexanderi
| `--+--Proterosuchus goweri
| `--Proterosuchus yuani
`--+*-Kalisuchus rewanensis
|*-Vonhuenia fredericki
|--Antarctanax shackletoni
`--+--Sarmatosuchus otschevi
`--+*-Avalonianus sanfordi
|*-Basutodon ferox
|*-"Cinizasaurus hunti"
|*?"Likhoelesaurus ingens"
|*-"Massospondylus" hislopi
|*-"Megalosaurus" cloacinus
|*-"Megalosaurus" obtusus
|*-Paleosaurus cylindrodon
|*-Paleosaurus? subcylindrodon
|*-Picrodon herveyi
|*-"Thecodontosaurus" elizae
|*-Wangisuchus tzeyii
|*?Zanclodon? bavaricus
|*?Zanclodon laevis
|*-Zatomus sarcophagus
|*-Protopyknosia
| |--Kranosaura kuttyi
| `--Triopticus primus
|--Cuyosuchus hueinei
`--+--Erythrosuchia
| |--Fugusuchus hejiapanensis
| `--+*-Uralosaurus magnus
| |--Guchengosuchus shiguaiensis
| `--+--+--Garjiania madiba
| | `--Garjiania prima
| `--+*-Shanshisuchus? kuyeheensis
| |--Erythrosuchus africanus
| `--+--Chalishevia cothurnata
| `--Shanshisuchus shanshisuchus
`--Eucrocopoda
|--Asperoris mnyama
|--Dorosuchus noetus
`--+--Euparkeriidae
| |--Euparkeria capensis
| `--Osmolskina czatkowicensis
`--+--Halazhaisuchus qiaoensis
`--+*-Aenigmaspina pantyffynonensis
|--Proterochampsia
| |--+--Litorosuchus somnii
| | `--Vancleavea campi
| `--+--Doswelliidae
| | |--Doswellia kaltenbachi
| | `--Jaxtasuchus salomani
| `--Proterochampsidae
| |--+--Proterochampsa barrionoevoi
| | `--Proterochampsa nodosa
| `--+--Cerritosaurus binsfeldi
| `--+--Tropidosuchus romeri
| `--Rhadinosuchinae
| |--Pseudochampsa ischigualastensis
| `--+--Gualosuchus reigi
| `--+--Chanaresuchus bonapartei
| `--Rhadinosuchus gracilis
`--Archosauria
|*-"Cryptoraptor lockleyi"
|*-Lametasaurus indicus
|*-Lukousaurus yini
|*-"Plateosaurus" ornatus
|*-Shansaraiki insafi
|*-Tanystrosuchus posthumus
|*-Vitakrisaurus saraiki
|--Pseudosuchia
| Crurotarsi
| |*-Dasygnathoides longidens
| |--Phytosauria
| | |--Diandongosuchus fuyuanensis
| | `--+--"Zanclodon" arenaceus
| | `--Parasuchidae
| | |*-Belodon plieningeri
| | |*?Centemodon sulcatus
| | |*-Clepsysaurus? fraserianus
| | |*-Clepsysaurus pennsylvanicus
| | |*-Clepsysaurus? veatleianus
| | |*-"Megalosaurus" terquemi
| | |*-Palaeoctanus appalachianus
| | |*-Palaeoctanus? aulacodus
| | |*-Palaeoctanus? dumblianus
| | |*-Palaeoctanus? orthodon
| | |*-"Paleosaurus" platyodon
| | |*-"Paleosaurus" stricklandi
| | |*-Phytosaurus cylindricodon
| | |*-Rileyasuchus bristolensis
| | |*-Rileyasuchus? stutchburyi
| | |*-Rutiodon? leaii
| | |*-Suchoprion cyphodon
| | |*-Suchoprion? sulcidens
| | |--Wannia scurriensis
| | `--+--+--Parasuchus bransoni
| | | `--+--Parasuchus angustifrons
| | | `--Parasuchus hislopi
| | `--+--"Paleorhinus" sawini
| | `--+--Ebrachosuchus neukami
| | `--+--"Paleorhinus" parvus
| | `--Mystriosuchinae
| | |--+--Rutiodon carolinensis (= Palaeonornis struthionoides)
| | | `--+--Angistorhinus talainti
| | | `--+--Angistorhinus alticephalus
| | | `--Angistorhinus grandis
| | ``+--"Smilosuchus" lithodendrorum
| | `--Leptosuchomorpha
| | |--+--Leptosuchus? studeri
| | | `--+--Leptosuchus crosbiensis
| | | `--+--Pravusuchus hortus
| | | `--+--Nicrosaurus kapffi
| | | `--+--Nicrosaurus? meyeri
| | | `--Coburgosuchus goekeli
| | `--+--Phytosaurus? doughtyi
| | `--+--Smilosuchus? adamanensis
| | `--+--Smilosuchus gregorii
| | `--+--Protome batalaria
| | `--+--Machaeroprosopus? andersoni
| | `--Mystriosuchini
| | |--Machaeroprosopus? jablonskiae
| | `--+--Machaeroprosopus? mccauleyi
| | `--+--+--Machaeroprosopus buceros
| | | `--Machaeroprosopus pristinus
| | `--+--Machaeroprosopus? zunii
| | `--+--"Machaeroprosopus" lottorum
| | `--+--+--Redondosuchus bermani
| | | `--Redondosuchus gregorii
| | `--+--Mystriosuchus steinbergeri
| | `--+--Mystriosuchus planirostris
| | `--Mystriosuchus westphali
| `--+--Nundasuchus songaensis
| `--Suchia
| |*-Barberenasuchus brasiliensis
| |--Ornithosuchia
| | |--Ornithosuchidae
| | | |--Ornithosuchus woodwardi
| | | `--+--Dynamosuchus collisensis
| | | `--+--Riojasuchus tenuisceps
| | | `--Venaticosuchus rusconii
| | `--+--Erpetosuchidae
| | | |--Parringtonia gracilis
| | | `--+--Dyoplax arenaceus
| | | `--+--Erpetosuchus granti
| | | `--+--Archeopelta arborensis
| | | |--Pagosovenator candelariensis
| | | `--Tarjadia ruthae
| | `--+--+--+--Revueltosaurus callenderi
| | | | `--Revueltosaurus hunti
| | | `--+--Acaenasuchus geoffreyi
| | | `--Euscolosuchus olseni
| | `--Aetosauria
| | |--Aetosauroides scagliai
| | `--Stagonolepididae
| | |--Stagonolepidoidea
| | | |--Stagonolepidinae
| | | | |--Polesinesuchus aurelioi
| | | | `--Stagonolepis robertsoni
| | | `--Desmatosuchinae
| | | |--"Stagonolepis" olenkae
| | | `--+--Neoaetosauroides engaeus
| | | `--+--+--Calyptosuchus wellesi
| | | | `--+--Adamanasuchus eisenhardtae
| | | | `--Scutarx deltatylus
| | | `--Desmatosuchini
| | | |--Gorgetosuchus pekinensis
| | | `--+--Longosuchus meadei
| | | `--+--Sierritasuchus macalpini
| | | `--+--Lucasuchus hunti
| | | `--+--Desmatosuchus smalli
| | | `--Desmatosuchus spurensis
| | `--Aetosaurinae
| | |--Stenomyti huangae
| | `--+--Aetosaurus ferratus
| | |--Coahomasuchus kahleorum
| | `--Typothoracinae
| | |--Apachesuchus heckerti
| | `--+--+--Redondasuchus rineharti
| | | `--Typothorax coccinarum
| | `--Paratypothoracini
| | |--Rioarribasuchus chamaensis
| | `--+--Tecovasuchus chatterjeei
| | `--Paratypothorax andressorum
| `--+--Gracilisuchidae
| | |--Gracilisuchus stipinicorum
| | `--+--Turfanosuchus dabanensis
| | `--Yonghesuchus sanbiensis
| `--+*-Arganasuchus dutuiti
| |*-Koilamasuchus gonzalezdiazi
| |--Ticinosuchus ferox
| `--Paracrocodylomorpha
| |--Mandasuchus tanyauchen
| |--Stagonosuchus nyassicus
| |--Poposauroidea
| | |--Youngosuchus sinensis
| | `--+--Qianosuchus mixtus
| | |*-Bromsgroveia walkeri
| | |--Ctenosauriscidae
| | | |--Arizonasaurus babbitti
| | | |--Ctenosauriscus koeneni
| | | |--Hypselorhachis mirabilis
| | | `--Xilousuchus sapingensis
| | `--+--Poposauridae
| | | |--Poposaurus gracilis
| | | `--Poposaurus langstoni
| | `--+--Lotosaurus adentus
| | `--Shuvosauridae
| | |--Sillosuchus longicervix
| | `--+--Shuvosaurus inexpectatus
| | `--Effigia okeeffeae
| `--Loricata
| |*-Decuriasuchus quartacolonia
| |*-Tikisuchus romeri
| |--Prestosuchidae
| | |--+--Prestosuchus chiniquensis
| | | `--Prestosuchus nyassicus
| | `--+--Luperosuchus fractus
| | `--Saurosuchus galilei
| `--+--Apatosuchus orbitoangulatus
| |--Etjosuchus recurvidens
| `--+--+--Batrachotomus kupferzellensis
| | `--Heptasuchus clarki
| `--+--Fasolasuchus tenax
| `--+--Teratosauroidea
| | Teratosauridae
| | |*-Vivaron haydeni
| | |--Rauisuchus tiradentes
| | `--+--Polonosuchus silesiacus
| | |--+--Postosuchus alisonae
| | | `--Postosuchus kirkpatricki
| | `--Teratosaurus suevicus
| `--Crocodylomorpha
| |*-Clarencea gracilis
| |*-Pedeticosaurus leviseuri
| |--Carnufex carolinensis
| |--Redondavenator quayensis
| `--+--Trialestidae
| | |--Pseudohesperosuchus jachaleri
| | `--Trialestes romeri
| `--+--Saltoposuchidae
| | |--Saltoposuchus connectens
| | `--+--Terrestrisuchus gracilis
| | `--Litargosuchus leptorhynchus
| `--+--+--Hesperosuchus agilis
| | `--+--Dromicosuchus grallator
| | `--Kayentasuchus walkeri
| `--+--Sphenosuchus acutus
| `--+--Dibothrosuchus elaphros
| `--+--Hallopodidae
| | |--Almadasuchus figarii
| | `--+--Hallopus victor
| | `--Macelognathus vagans
| `--+--Jungarrsuchus sloani
| |*-+--Eopneumatosuchus colberti
| | `--Thalattosuchia
| | |--Teleosauroidea
| | | |*-Cystosaurus
| | | |*-"Megalosaurus" mersensis
| | | |--Plagiophthalmosuchus gracilirostris
| | | `--+--Clovesuurdameredeor stephani
| | | |--Macrospondylus bollensis
| | | |--Teleosauridae
| | | | |--Indosinosuchus potamosiamensis
| | | | |--Mystriosaurus laurillardi
| | | | |--Teleosaurinae
| | | | | |--Platysuchus multiscrobiculatus
| | | | | `--Teleosaurus cadomensis
| | | | `--Aeolodontinae
| | | | |--Mycterosuchus nasutus
| | | | `--+--Aeolodon priscus
| | | | `--+--Bathysuchus megarhinus
| | | | `--Sericodon jugleri
| | | `--Machimosauridae (Steneosauridae)
| | | |--Charitomenosuchus leedsi
| | | |--Seldsienean megistorhynchus
| | | `--+--Deslongchampsina larteti
| | | `--Machimosaurinae (Steneosaurinae)
| | | |--+--Proexochokefalos bouchardi
| | | | `--Proexochokefalos heberti
| | | |--Andrianavoay baroni
| | | |--Steneosaurus rostromajor
| | | |--Lemmysuchus obtusidens
| | | |--Yvridiosuchus boutilieri
| | | `--Machimosaurini
| | | |--Machimosaurus buffetauti
| | | |--Machimosaurus mosae
| | | `--+--Machimosaurus hugii
| | | `--Machimosaurus rex
| | `--Metriorhynchoidea
| | |--Pelagosaurus typus
| | `--+--Peipehsuchus teleorhinus
| | `--+--Teleidosaurus calvadosii
| | `--+--+--Magyarosuchus fitosi
| | | `--Opisuchus meieri
| | `--+--+--Eoneustes bathonicus
| | | `--Eoneustes gaudryi
| | `--+--Zoneait nargorum
| | `--Metriorhynchidae
| | |*-Enaliosuchus macrospondylus
| | |--Metriorhynchinae
| | | |--Thallatosuchus superciliosus
| | | `--+--+--Gracilineustes acutus
| | | | `--Gracilineustes leedsi
| | | `--+--+--Maledictosuchus nuyivijanan
| | | | `--Maledictosuchus riclaensis
| | | `--+--Metriorhynchus brevirostris
| | | `--Rhacheosaurini
| | | |--+--"Cricosaurus" saltillensis
| | | | `--"Metriorhynchus" palpebrosus
| | | `--+--Rhacheosaurus gracilis (= Megalosaurus gracilis)
| | | `--+--+*-Cricosaurus puelchorum
| | | | |--Cricosaurus bambergensis
| | | | `--+--Cricosaurus elegans
| | | | `--Cricosaurus suevicus
| | | `--+--Cricosaurus lithographicus
| | | |--Cricosaurus araucanensis
| | | `--+--Cricosaurus schroederi
| | | `--Cricosaurus vignaudi
| | `--Geosaurinae
| | |--+--"Metriorhynchus" casamiquelai
| | | `--"Metriorhynchus" westermanni
| | `--+--Neptunidraco ammoniticus
| | `--+--"Metriorhynchus" brachyrhynchus
| | `--Geosaurini
| | |--+--Tyrannoneustes lythrodectikos
| | | `--+--Purranisaurus potens
| | | `--+--Torvoneustes coryphaeus
| | | `--+--Torvoneustes carpenteri
| | | `--Torvoneustes mexicanus
| | `--+--Geosaurina
| | | |--Ieldraan melkshamensis
| | | `--+--Geosaurus grandis
| | | `--+--Geosaurus giganteus
| | | `--Geosaurus lapparenti
| | `--+--Plesiosuchina
| | | |--Plesiosuchus manselii
| | | `--Suchodus durobrivensis
| | `--"Dakosaurina"
| | |?-"Dakosaurus" lissocephalus
| | |--Dakosaurus andiniensis
| | |--Dakosaurus maximus (= Megalosaurus gracilis, ?= Megalosaurus schnaitheimii)
| | `--Dakosaurus nicaeensis (= Aggiosaurus nicaeensis)
| `--Crocodyliformes
| |*-Karatausuchus sharovi
| |*-Kemkemia auditorei
| |*-"Maroccanoraptor elbegiensis"*
| |--Protosuchia
| | Protosuchidae
| | |--Coloradisuchus abelini
| | |--Hemiprotosuchus leali
| | |--Hoplosuchus kayi
| | |--Orthosuchus stormbergi
| | |--Edentosuchus tienshanensis
| | `--+--Protosuchus haughtoni
| | `--Protosuchus richardsoni
| |--Gobiosuchidae
| | |--Cassissuchus sanziuami
| | |--Lisboasaurus estesi
| | `--+--Gobiosuchus kielanae
| | `--Zaraasuchus shepardi
| `--Mesoeucrocodylia
| |*-Lusitanisuchus mitracostatus
| `--+--Shartegosuchoidea
| | |--+--Zosuchus davidsoni
| | | `--+--+--Sichuanosuchus huidongensis
| | | | `--Sichuanosuchus shuhanensis
| | | `--+--Shantungosuchus chuhsienensis
| | | `--Neuquensuchus universitas
| | `---Shartegosuchidae
| | |*-Kyasuchus saevi
| | |--Nominosuchus matutinus
| | `--+--Fruitachampsa callisoni
| | `--+--Adzosuchus fuscus
| | `--Shartegosuchus asperopalatum
| `--+--Hsisosuchidae
| | |--Hsisosuchus chungkingensis
| | `--+--Hsisosuchus chowi
| | `--Hsisosuchus dashanpuensis
| `--Metasuchia
| |--Microsuchus schilleri
| |--Notosuchia
| | Notosuchiformes
| | |*-Brasileosaurus pachecoi
| | |--+?*Cunampaia simplex
| | | |--Mahajangasuchidae
| | | | |--Kaprosuchus saharicus
| | | | `--Mahajangasuchus insignis
| | | `--Peirosauridae (Trematochampsidae)
| | | |*?Fortignathus felixi
| | | |--Amargasuchus minor
| | | `--+--Miadanasuchus oblita
| | | `--+?-Stolokrosuchus lapparenti
| | | `--+--Pepesuchinae
| | | | |*-Roxochampsa paulistanus
| | | | |--Caririsuchus camposi
| | | | `--+--Itasuchus jesuinoi
| | | | `--+--Barreirosuchus franciscoi
| | | | `--Pepesuchus deiseae
| | | `--Peirosaurinae (Trematochampsinae)
| | | |--Hamadasuchus rebouli
| | | `--+*-Bayomesasuchus hernandezi
| | | |--Lomasuchus palpebrosus
| | | `--+--+--Barcinosuchus gradilis
| | | | `--Gasparinisuchus peirosauroides
| | | `--+--Montealtosuchus arrudacamposi
| | | |--Trematochampsa taqueti
| | | |--Peirosaurus torminni
| | | `--Uberabasuchus terrificus
| | |--Uruguaysuchidae
| | | |--Araripesuchus tsangatsangana
| | | |--+--Anatosuchus minor
| | | | `--Araripesuchus wegeneri
| | | `--+--Araripesuchus buitreaensis
| | | `--+--Araripesuchus gomesii
| | | `--+--Araripesuchus patagonicus
| | | `--Uruguaysuchus aznarezi
| | `--Ziphosuchia
| | |--Libycosuchus brevirostris
| | `--+--Simosuchus clarki
| | `--+--Candidodontidae
| | | |--Lavocatochampsa sigogneaurussellae
| | | `--+--Candidodon itapecuruensis
| | | `--+--Malawisuchus mwakasyungutiensis
| | | `--Pakasuchus kapilimai
| | `-----Gondwanasuchia
| | |--Sphagesauria
| | | |*-Coringasuchus anisodontis
| | | |*-Coronelsuchus civali
| | | |--Notosuchidae
| | | | |--Llanosuchus tamaensis
| | | | `--Notosuchus terrestris
| | | `--+--Morrinhosuchus luziae
| | | `--+--Labidiosuchus amicum
| | | `--+--+--Mariliasuchus amarali
| | | | `--Mariliasuchus robustus
| | | `--Sphagesauridae
| | | |--+--Adamantinasuchus navae
| | | | `--Yacarrerani boliviensis
| | | `--+--Caipirasuchinae
| | | | |--+--Caipirasuchus mineirus
| | | | | `--+--Caipirasuchus attenboroughi
| | | | | `--Caipirasuchus stenognathus
| | | | `--+--Caipirasuchus montealtensis
| | | | `--Caipirasuchus paulistanus
| | | `--Sphagesaurinae
| | | |--Sphagesaurus heunei
| | | `--+--Caryonosuchus pricei
| | | `--Armadillosuchus arrudai
| | `--+--Chimaerasuchus paradoxus
| | `--+*-Razanandrongobe sakalavae
| | |--Comahuesuchus brachybuccalis
| | `--Sebecosuchia
| | |--Pehuenchesuchus enderi
| | `--+--+--Doratodon carcharidens
| | | `--Doratodon ibericus
| | `--Sebecoidea
| | |*-Ilchunaia parca
| | |?-+--Sahitisuchus fluminensis
| | | `--+--Iberosuchidae
| | | | |--Bergisuchus dietrichbergi
| | | | `--+--Iberosuchus macrodon
| | | | `--Iberosuchus crassiproratus
| | | `--Sebecidae
| | | |--Lorosuchus nodosus
| | | `--+--+--Ayllusuchus fernandezi
| | | | `--Bretesuchus bonapartei
| | | `--+--Barinasuchus arveloi
| | | |--Eremosuchus elkoholicus
| | | |--Ogresuchus furatus
| | | `--+--Langstonia huilensis
| | | |--Sebecus icaeorhinus
| | | `--Zulmasuchus querejazus
| | `--Baurusuchia
| | Baurusuchidae
| | |--Cynodontosuchus rothi
| | `--+--Gondwanasuchus scabrosus
| | |--Pabwehshi pakistanensis (= syntype of "Vitakridrinda sulaimani")
| | `--+--Pissarrachampsinae
| | | |--Campinasuchus dinizi
| | | `--+--Pissarrachampsa sera
| | | `--Wargosuchus australis
| | `--Baurusuchinae
| | |--Aphaurosuchus escharafacies
| | `--+--Aplestosuchus sordidus
| | `--+--Stratiotosuchus maxhechti
| | `--+--Baurusuchus albertoi
| | `--+--Baurusuchus pachecoi
| | `--Baurusuchus salgadoensis
| `--Neosuchia
| |*-Montsecosuchus depereti
| |--+--Paralligatoridae
| | | |*-Paralligator? sungaricus
| | | |--Rugosuchus nonganensis
| | | `--+--Batrachomimus pastobonensis
| | | `--+--Shamosuchus djadochtaensis
| | | `--+--Tarsomordeo winkleri
| | | `--+--Kansajsuchus extensus
| | | `--+--Paralligator major
| | | `--+--Paralligator gradilifrons
| | | `--Scolomastax salhsteini
| | |--Wannchampsus kirpachi
| | `--Atoposauridae
| | |--Aprosuchus ghirai
| | |--+--Brillanceausuchus babouriensis
| | | `--+--Sabresuchus ibericus
| | | `--Sabresuchus sympiestodon | |
| | |--+--Theriosuchus? grandinaris
| | | `--+--Theriosuchus pusillus
| | | `--+--Knoetschkesuchus guimarotae
| | | `--Knoetschkesuchus langenbergensis
| | `--+--Alligatorium meyeri
| | |--"Alligatorium" franconicum
| | `--+--+--Alligatorellus bavaricus
| | | `--Alligatorellus beaumonti
| | `--+--Atoposaurus jourdani
| | `--Atoposaurus oberndorferi
| |--Bernissartiidae
| | |--Bernissartia fagessi
| | `--Koumpiodontosuchus aprosdokitii
| |--Khoratosuchus jintasakuli
| |--Goniopholididae
| | |--Calsoyasuchus valliceps
| | |--"Goniopholis" stovalli
| | `--+*-Sunosuchus? phuwiangensis
| | |--Eutretauranosuchus delfsi
| | |--Sunosuchus? junggarensis
| | `--+--Sunosuchus miaoi
| | `--+--Siamosuchus phuphokensis
| | `--+--Amphicotylus lucasii
| | `--+*-Denazinosuichus kirtlandicus
| | |--+--Nannosuchus gracilidens
| | | `--+--Goniopholis baryglyphaeus
| | | `--+--Goniopholis kiplingi
| | | `--Goniopholis simus
| | `--+--Hulkepholis willetti
| | `--+--+--Hulkepholis? plotos
| | | `--Hulkepholis? rori
| | `--+--Anteophthalmosuchus escuchae
| | `--+--Anteophthalmosuchus epikrator
| | `--Anteophthalmosuchus hooleyi
| |--Paluxysuchidae
| | |--Deltasuchus motherali
| | `--Paluxysuchus newmani
| |--Coelognathosuchia
| | Tethysuchia
| | |--Pholidosauridae
| | | |--Meridiosaurus vallisparidisi
| | | `--+*-Dakotasuchus kingi
| | | |--+--Pholidosaurus purbeckensis
| | | | `--Pholidosaurus schaumbergensis
| | | `--+--+--Oceanosuchus boencensis
| | | | |--Sabinosuchus coahuilensis
| | | | `--Woodbinesuchus byersmauricei
| | | `--+--+--Terminonaris browni
| | | | `--Terminonaris robusta
| | | `--+--Chalawan thailandicus
| | | |--Sarcosuchus hartti
| | | `--Sarcosuchus imperator
| | `--Dyrosauroidea
| | |--Vectisuchus leptognathus
| | `--+--Elosuchidae
| | | |--Elosuchus broinae
| | | `--Elosuchus cherifiensis
| | `--Dyrosauridae
| | |--Chenanisuchus lateroculi
| | `--+--Anthracosuchus balrogus
| | `--+--Cerrejonisuchus improcerus
| | `--+--Phosphatosaurinae
| | | |--Sokotosuchus ianwilsoni
| | | `--Phosphatosaurus gavialoides
| | `--+--Rodeosuchus machukiru
| | `--Hyposaurinae (Dyrosaurinae)
| | |--Dorbignysuchus niatu
| | `--+--+--Hyposaurus derbianus
| | | |--Hyposaurus natator
| | | `--Hyposaurus rogersii
| | `--+--+--Arambourgisuchus khouribgaensis
| | | `--+--Luciasuchus lurusinga
| | `--+--+--Dyrosaurus maghribensis
| | | `--Dyrosaurus phosphaticus
| | `--+--Acherontisuchus guajiraensis
| | `--+--Atlantosuchus coupatezi
| | `--+*-Rhabdognathus rarus
| | |--Rhabdognathus? aslerensis
| | `--+--Rhabdognathus? keiniensis
| | `--+--Congosaurus compressus
| | `--Congosaurus bequaerti
| |--Stomatosuchidae
| | |--+--Laganosuchus thaumastos
| | | `--Laganosuchus maghrebensis
| | `--Stomatosuchus inermis
| |--Susisuchidae
| | |--+--Susisuchus anatoceps
| | | `--Susisuchus jaguaribensis
| | `--+--Isisfordia duncani
| | `--Isisfordia selaslophensis
| `--Eusuchia
| |--Hylaeochampsidae
| | |?-Pachycheilosuchus trinquei
| | `--+--Pietraroiasuchus ormezzanoi
| | `--+--Turcosuchus okani
| | `--+--Unasuchus reginae
| | `--+--Acynodon iberoccitanus
| | `--+--Acynodon? adriaticus
| | `--+--Hylaeochampsa vectiana
| | `--Iharkutosuchus makadii
| |--Allodaposuchidae
| | |--Arenysuchus gascabadiolorum
| | `--+--+--Allodaposuchus iberoamericanus
| | | `--Allodaposuchus precedens
| | `--+--Allodaposuchus subjuniperus
| | `--+--+--Allodaposuchus hulki
| | | `--Allodaposuchus palustris
| | `--+--Allodaposuchus fontisensis
| | `--+--Allodaposuchus mechinorum
| | `--Allodaposuchus megadontos
| `--+--Portugalosuchus azenhae
| `--+--"Borealosuchidae"
| | |--Borealosuchus sternbegii
| | `--+--Borealosuchus formidabilis
| | `--+--Borealosuchus acutidentatus
| | |--Borealosuchus threeensis
| | `--Borealosuchus wilsoni
| `--+*-Pristichampsus rollinati
| |--Planocraniidae
| | |--Planocrania? hengdongensis
| | `--+--Planocrania datangensis
| | `--+--Boverisuchus magnifrons
| | `--Boverisuchus vorax
| `--Crocodylia
| |--Alligatoroidea
| | |--Leidyosuchus canadensis
| | `--+--+--Deinosuchus hatcheri
| | | |--Deinosuchus riograndensis
| | | `--Deinosuchus schwimmeri
| | `--+--Diplocynodontinae
| | | |--Diplocynodon darwini
| | | |--Diplocynodon deponiae
| | | `--+--Diplocynodon muelleri
| | | |--Diplocynodon ratelii
| | | |--Diplocynodon tormis
| | | `--+--Diplocynodon remensis
| | | `--+--Diplocynodon hantoniensis
| | | `--+--Diplocynodon elavericus
| | | `--Diplocynodon ungeri
| | `--Globidonta
| | |--+--Brachychampsa montana
| | | `--Brachychampsa sealeyi
| | `--+--Albertochampsa langstoni
| | |--Stangerochampsa mccabei
| | `--Alligatoridae
| | |--Alligatorinae
| | | |--Ceratosuchus burdoshi
| | | |--Wannaganosuchus brachymanus
| | | |--Navajosuchus mooki
| | | |--+--Allognathosuchus polyodon
| | | | |--Allognathosuchus wartheni
| | | | `--+--Procaimanoidea? kayi
| | | | `--+--Arambourgia gaudryi
| | | | `--Procaimanoidea utahensis
| | | `--+--Alligator prenasalis
| | | `--+--Alligator mcgrewi
| | | `--+--Alligator sinensis
| | | `--+--Alligator mefferdi
| | | `--Alligator mississippiensis
| | `--Caimaninae
| | |--Culebrasuchus mesoamericanus
| | `--+--Gnatusuchus pebasensis
| | `--+--Globidontosuchus brachyrostris
| | `--+--+--Eocaiman palaeocenicus
| | | `--+--Eocaiman cavernensis
| | | `--Eocaiman itaboraiensis
| | `--+--Kuttanacaiman iquitosensis
| | `--+--+--+--Bottosaurus harlani
| | | | `--Tsoabichi greenriverensis
| | | `--+--Paleosuchus palpebrosus
| | | `--Paleosuchus trigonatus
| | `--+--+--"Caiman" gasparinae
| | | `--+--+--Mourasuchus amazonensis
| | | | `--Mourasuchus pattersoni
| | | `--+--Mourasuchus arendsi
| | | `--Mourasuchus atopus
| | `--+*-Centenariosuchus gilmorei
| | |*-Necrosuchus ionensis
| | |--+--Acresuchus pachytemporalis
| | | `--+--Purussaurus brasiliensis
| | | |--Purussaurus mirandai
| | | |--Purussaurus amazonicus
| | | `--Purussaurus? terror (= Dinosuchus terror)
| | `--+--"Caiman" wannlangstoni
| | `--Jacarea
| | |*-Caiman? brevirostris
| | |--Melanosuchus niger
| | `--+--Caiman latirostris
| | `--+--Caiman crocodilus
| | `--Caiman yacare
| `--+--Asiatosuchus germanicus
| |--Prodiplocynodon langi
| `--+--+--"Crocodilus" affinis
| | `--"Crocodilus" depressifrons
| `--+--+--"Crocodilus" acer
| | `--Brachyuranochampsa eversolei
| `--Longirostres
| |--Gavialoidea
| `--Crocodyloidea
`--Pan-Aves
|--Aphanosauria
| |*?Arctosaurus osborni
| |--Spondylosoma absconditum
| `--+--Dongusuchus efremovi
| |--Teleocrater rhadinus
| `--Yarasuchus deccanensis
`--Avifilopluma
Ornithodira
|*?Avipes dillstedtianus
|--Pterosauromorpha
| |--Maehary bonapartei
| |?-Protoavis texensis (in part)
| `--+--Scleromochlus taylori
| |--Lagerpetidae
| | |*-Faxinalipterus minimus
| | |*-Kongonaphon kely
| | |--Lagerpeton chanerensis
| | `--+--Ixalerpeton polesinensis
| | `--+--Dromomeron gregorii
| | `--+--Dromomeron romeri
| | `--Dromomeron gigas
| `--Pterosauria
| |--Rhaeticodactylidae
| | |--+--Carniadactylus
| | | `--+--Caviramus
| | | `--Raeticodactylus
| | `--+--Arcticodactylus
| | |--Austriadraco
| | `--Seazzadactylus
| `--+--Preondactylia
| | |--Austriadactlyus
| | `--Preondactylus
| `--+--Peteinosaurus
| `--Macronychoptera
| |--Dimorphodontia
| `--+--Eudimorphodon
| `--Novialoidea
| |--Campylognathoididae
| `--Breviquartossa
| |*-"Archaeopteryx" "vicensensis"
| |*-Laopteryx prisca
| |--Rhamphorhynchidae
| | |*-"Crateropteryx colei"
| | |--Scaphognathinae
| | `--Rhamphorhynchinae
| | `--Rhamphorhynchus muensteri
| `--Pterodactylomorpha
| |--Sordes
| `--Monofenestrata
| |--Darwinoptera
| | |--Pterorhynchus
| | `--Wukongopteridae
| | |--Kunpengopterus
| | `--Wukongopterinae
| `--Pterodactyliformes
| |--Changchengopterus
| `--Caelidracones
| |*-Eurolimnornis corneti
| |--Anurognathidae
| | |--"Dimorphodon" weintraubi
| | `--+--Batrachognathinae
| | `--Anurognathinae
| `--Pterodactyloidea
| |--Kryptodracon
| `--Lophocratia
| |*-Herbstosaurus pigmaeus
| |--Archaeopterydactyloidea
| | |--Germanodactylidae
| | `--Euctenochasmatia
| | |--Pterodactylidae
| | | `--Pterodactylus antiquus
| | `--+--Ardeadactylus
| | `--Ctenochasmatoidea
| | |--Gallodactylidae
| | `--Ctenochasmatidae
| | |--Ctenochasmatinae
| | | |--Ctenochasma
| | | `--Pterodaustrini
| | `--+--Kepodactylus
| | `--+--Elanodactylus
| | `--+--Gnathosaurinae
| | `--Moganopterinae
| `--Eupterodactyloidea
| |*-Cimoliornis diomedeus
| |--Pteranodontia
| | |--Nyctosauridae
| | | `--Cretornis hlavaci
| | `--+--Alamodactylus
| | `--Ornithocheiroidea
| | |*-Piksi barbarulna
| | |--Pteranodontidae
| | `--Ornithocheiromorpha
| | Lanceodontia
| | |--Istiodactyliformes
| | | |--Lonchodectidae
| | | `--+--Haopterus
| | | |--Linglongopterus
| | | |--Mimodactylus
| | | `--+--Hongshanopterus
| | | `--Istiodactylidae
| | | |--Nurhachius
| | | `--Istiodactylinae
| | `--Anhangueria
| | |--Boreopteridae
| | `--Ornithocheirae
| | |--Ornithocheiridae
| | `--+--Targaryendraconia
| | | |--Cimoliopteridae
| | | `--Targaryendraconidae
| | `--+--Hamipteridae
| | `--Anhangueridae
| | |--Tropeognathinae
| | `--+--Coloborhynchinae
| | `--Anhanguerinae
| `--Azhdarchoidea
| |*-"Palaeornis" cliftii
| |--Tapejaromorpha
| | |--Bennettazhia
| | |--Caupedactylia
| | `--Tapejaridae
| | |--Sinopterinae
| | `--Tapejarinae
| `--Neoazhdarchia
| |--Dsungaripteromorpha
| | |--Dsungaripteridae
| | | |--Noripterinae
| | | `--Dsungaripterinae
| | `--Thalassodrominae
| `--Neopterodactyloidea
| |--Chaoyangopteridae
| | |--Eoazhdarcho
| | `--Chaoyangopterinae
| `--+--Radiodactylus
| `--Azhdarchidae
| |*-Aralazhdarcho bostobensis (?= Samrukia nessovi)
| |*?Palaeocursornis corneti
| |--Azhdarcho
| `--Quetzalcoatlinae
`--Dinosauromorpha
Dinosauriformes
|--Lagosuchus talampayensis
|*-Nyasasaurus parringtoni
|*-Saltopus elginensis
`--Dracohors
Avipluma
Dinosauria
|*-Apatodon mirus
|--Ornithischia
| |--Lewisuchus admixtus
| `--+--Soumyasaurus aenigmaticus
| `--+*-Gamatavus antiquus
| |--Asilisaurus kongwe
| `--Sulcimentisauria
| |--Diodorus scytobrachion
| `--+--Technosaurus smalli
| `--Parapredentata
| |--Silesauridae
| | |--Ignotosaurus fragilis
| | `--Silesaurus opolensis
| |--Sacisaurus agudoensis
| `--+--+--Lutungutali sitwensis
| | `--Kwanasaurus williamparkeri
| `--+--Eucoelophysis baldwini
| `--+--Pisanosaurus mertii
| `--+*?Thecospondylus horneri
| |*?Tichosteus? aequifacies
| |*?Tichosteus lucasanus
| |--Laquintasaura venezuelae
| `--+--Lesothosaurus diagnosticus
| `--Genasauria
| |--Thyreophora
| | |*-Regnosaurus northamptoni
| | |--Scutellosaurus colberti
| | `--+--Emausaurus ernsti
| | `--+--Scelidosaurus harrisonii
| | `--Eurypoda
| | |--Stegosauria
| | | |--"Astrodon" pusillus
| | | |--Chialingosaurus kuani
| | | `--Craterosaurus pottonensis
| | `--Ankylosauria
| | |--Sarcolestes leedsi
| | `--Struthiosaurus austriacus
| `--Neornithischia
| |--Eocursor parvus
| `--+--+--Agilisaurus louderbacki
| | `--Hexinlusaurus multidens
| `--Cerapoda
| |--Ornithopoda
| | |--Nanosaurus agilis
| | `--+--Changchunsaurus parvus
| | `--+--Jeholosauridae
| | `--+--Haya griva
| | `--+--Orodromeus makelai
| | `--+--Kulindadromeus zabaikalicus (= Lepidocheirosaurus natatilis)
| | `--Clypeodonta
| | |--Hypsilophodontidae
| | `--+--Thescelosauridae
| | `--+--+--Convolosaurus marri
| | | `--Yueosaurus tiantaiensis
| | `--Iguanodontia
| | |--Tenontosaurus spp.
| | `--+--Rhabdodontomorpha
| | `--+--+--Eousdryosaurus nanohallucis
| | | `--Elasmaria
| | | |--Fulgurotherium australe
| | | `?-Loncosaurus argentinus
| | `--Dryomorpha
| | |--Dryosauridae
| | `--Ankylopollexia
| | |--Camptosauridae
| | | `--Camptosaurus dispar (?= Symphyrophus musculosus)
| | `--Styracosterna
| | |*-"Cetiosaurus" brachyurus
| | |--Theiophytalia kerri
| | `--Neoiguanodontia
| | |--Hypselospinus fittoni
| | `--Hadrosauriformes
| | |--Iguanodontidae
| | `--Hadrosauroidea
| | |--Probactrosaurus gobiensis
| | `--Hadrosauromorpha
| | |--Parrosaurus missouriensis
| | `--+--Bactrosaurus johnsoni
| | |--"Troodon" isfarensis
| | `--+--Telmatosaurus transsylvanicus (Elopteryx nopscai referred)
| | |--Hypsibema crassicauda
| | `--Hadrosauridae
| `--Marginocephalia
| |--Pachycephalosauria
| | |--Heterodontosauridae
| | | |--Fruitadens haagarorum (= cf. Coelurosaurus)
| | | `--Heterodontosaurinae
| | `--+--Tianyulong confuciusi
| | `--+--Echinodon becklesii
| | `--+--Wannanosaurus yansiensis
| | `--+--Goyocephale lattimorei
| | `--Pachycephalosauridae
| `--Ceratopsia
| |--+--Chaoyangosauridae
| | `--Psittacosauridae
| `--Neoceratopasia
`--Saurischia
|*-Alwalkeria maleriensis (in part)
|*-Herrerasauria
| Herrerasauridae
| |--Staurikosaurus pricei
| `--Herrerasaurinae
| |--Gnathovorax cabreirai
| |--Sanjuansaurus gordilloi
| `--Herrerasaurus ischigualastensis
|*-+--Chindesaurus bryansmalli
| `--Tawa hallae
|*-Daemonosaurus chauliodus
|*-Eoraptor lunensis
|*-Eodromaeus murphi
`--Eusaurischia
|--Sauropodomorpha
`--Theropoda